1) There are at least 8 *independent* indications that Pleistocene archaic Homo (H.erectus cs) was semi-aquatic:
• Archaic Homo's atypical tooth-wear was caused by "sand and oral processing of marine mollusks", Towle cs 2022 https://onlinelibrary.wiley.com/doi/10.1002/ajpa.24500
• H.erectus s.s. fossilized typically (always?) in coastal sediments, e.g. Mojokerto amid barnacles & corals, Trinil amid Pseudodon & Elongaria, Sangiran-17 in "brackish marsh near the coast".
• Stephen Munro discovered sea-shell engravings made by H.erectus, Joordens cs 2015 Nature 518:228–231 https://pubmed.ncbi.nlm.nih.gov/25470048/
• Ear exostoses (H.erectus & H.neand.) develop after years of cold(er) water irrigation https://www.ncbi.nlm.nih.gov/pmc/articles/PMC5696936/
• Pachyosteosclerosis is typically & exclusively seen in slow+shallow-diving tetrapods (de Buffrénil cs 2010 J.Mamm.Evol.17:101-120), e.g. erectus’ parietal bone is 2x as thick as in gorillas!
• Brain size in erectus (2x apes-australopiths) is facilitated by sea-food, e.g. DHA docosahexaenoic acid in shellfish etc., e.g. Odontocetes, Pinnipedia.
• Pleistocene Homo even colonized overseas islands (Flores & later even Luzon) https://www.academia.edu/36193382/Coastal_Dispersal_of_Pleistocene_Homo_2018
• Homo’s stone tool use & dexterity is typical for molluscivores, cf. sea-otters etc.

2) There's also no doubt that Mio-Pliocene Hominoidea were already "bipedal" = aquarboreal (aqua=water, arbor=tree) = regular BP wading + vertic.climbing + below-branch-hanging in swamp/coastal forests
https://www.gondwanatalks.com/l/the-waterside-hypothesis-wading-led-to-upright-walking-in-early-humans/
https://www.sciencedirect.com/science/article/abs/pii/S0169534702024904

3) IMO it's clear: afarensis-->boisei = fossil Gorilla // africanus-->robustus = fossil Pan (but I rarely get support & even less opposition to this).
https://www.academia.edu/8732353/Marc_Verhaegens_papers_in_Human_Evolution

4) IMO (my 2022 book p.299), Plate Tectonics played an essential role in hominoid splittings, but how *exactly* is still uncertain, e.g.
• India approaching Eurasia: Old World monkey/ape (some Catarrhini reaching these islands became Hominoidea),
• India further under Eurasia: lesser/great ape split (E/W),
• Mesopotamian Seaway closure c 15 Ma (mill.yrs ago): hominids/pongids (W/E), https://www.nature.com/articles/s41598-019-45308-7 + comments
• Red Sea fm, colonized by hominids s.s. (Homo-Pan-Gorilla),
• N-Rift fm (colonized by Gorilla-Praeanthropus: Lucy etc.,
• Red Sea opening into Gulf 6–5 Ma: Homo-->S.Asia, Pan-->E.Africa,
• S-Rift fm, colonized by Pan-Australopithecus: Taung etc.

5) More speculative: island evolution = drastic innovations? esp.
-- India approaching S-Eurasia 30?25 Ma: island archipel fm:
(some?) Catarrhini reaching these islands + coastal forests underwent drastic changes vs "monkeys":
larger body & brain, very broad body=pelvis/thorax/esp.sternum + lateral scapulas = lateral leg & arm (+upward) movements, centrally-placed spine (vs dorsally- in most mammals), tail loss...
-- "Sunda hypothesis": some hominoids in evolved into Homo s.s. = shellfish-diving = "aquatic"ape" s.s.??

These new insights in PA require a thorough re-arrangement of the old paleo-anthropology,
but many (most??) PAs still believe their ancestors ran after antelopes over African savannas (the endurance-running fantasy)!! :-D

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