• Which Mammals and Birds Survived the Great K-T Extinction, and Why?

    From Peter Nyikos@21:1/5 to All on Wed Nov 9 18:14:00 2022
    The Great K-T extinction spelled the doom of the giant sea reptiles, all dinosaurs
    (unless you count birds as dinosaurs), and many other groups of vertebrates and invertebrates.
    In fact, the damage caused by the asteroid that hit Yucatan was so great
    that no animal over 50 kilos is known to have survived.

    But some groups of birds and mammals did survive, including the remote ancestors
    of the four groups that have survived to this day: the neornithine birds and three groups of mammals:
    the placentals, the marsupials, and the monotremes.

    To keep this OP reasonably short, I will deal with the two most familiar of the four, the placentals and the modern birds.

    A 2013 _Science_ article [1] led to the general acceptance of the hypothesis that all living placentals
    have descended from a common ancestor that lived after the great K-T disaster.

    However, there are closely related eutherians [2] that also survived for a while, among
    whose descendants were the pantodonts, including *Coryphodon*, which was a bit larger than a tapir; and the tillodonts and taeniodonts. These groups died out during the Eocene, which ended almost halfway between the K-T disaster and our time.

    The case of birds is generally believed to be quite different: EVERY bird, living
    or extinct, that survived the K-T disaster is believed to be descended from the
    last common ancestor of living birds. However, there are some articles which hypothesize that there is one exception, a group of birds called the lithornids.
    More about them in my next post to this thread, some time tomorrow.

    [1] Maureen A. O'Leary et. al., "The Placental Mammal Ancestor and the Post–K-Pg Radiation of Placentals". Science. 339 (6120): 662–667.

    [2] Eutherians are the mammals, extant and extinct, that are more closely related to placentals than to marsupials.
    Similarly, metatherians are the mammals that are more closely related to marsupials than to placentals.


    Peter Nyikos
    Professor, Dept. of Mathematics -- standard disclaimer--
    University of South Carolina
    http://people.math.sc.edu/nyikos

    PS The "Why?" at the end of the thread title is two-edged: why did some survive,
    and why did others, including some similar-seeming ones, perish?

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From Peter Nyikos@21:1/5 to All on Thu Nov 10 13:07:07 2022
    On Wednesday, November 9, 2022 at 9:14:07 PM UTC-5, Peter Nyikos wrote:

    ...about a belief that is widespread among paleontologists and systematists, but with some dissenting articles:

    EVERY bird, living or extinct, that survived the K-T disaster is believed
    to be descended from the last common ancestor of living birds.
    However, there are some articles which hypothesize that there is one exception,
    a group of [long extinct] birds called the lithornids.

    The bone of contention here is: did lithornids descend from the
    last common ancestor (LCA) of living birds, or not -- did they split off
    from the branch leading to the LCA earlier?

    A bit of terminology is in order here before going on.

    Nowadays, it seems like the word "Aves", which once encompassed every bird
    in the classical sense, including *Archaeopteryx*, is coming to be confined to the birds
    that descended from the LCA of living birds. The usual name in the past for this much more
    restricted group is "Neornithes," and that is what is talked about below,
    in a paper by Livezey and Zusi, Zoological Journal of the Linnean Society, 2007, 149, 1–95.

    "A minor point of contention is the position of Lithornis (Houde, 1988), a relative to palaeognathous Neornithes, inferred to be the sistergroup of Tinamidae by Clarke & Norell (2002) and Clarke (2004), but inferred to be the sister-group of Neornithes
    by Clarke & Chiappe (2001), Leonard et al.(2005) and the present analysis (Fig. 12)."
    https://repository.si.edu/handle/10088/21644

    The first set of papers thus has lithornids inside Neornithes, the second set outside.


    By the way, lithornids are a very important group in the evolution of birds.

    I'll say more about that in my next post to this thread, to be done not long after I
    see that this one was posted.


    Peter Nyikos
    Professor, Dept. of Mathematics -- standard disclaimer--
    Univ. of South Carolina at Columbia
    http://people.math.sc.edu/nyikos

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From Peter Nyikos@21:1/5 to All on Thu Nov 10 14:20:10 2022
    The lithornids, about which I wrote in the preceding post to this thread,
    got their name (literally, "stone birds") from the fact that the first one
    to be described, *Lithornis,* was among the very first fossil birds
    to be described in the scientific literature.

    Owen described it in 1840, long before anyone had heard of Archaeopteryx.
    He thought it was a vulture.

    https://en.wikipedia.org/wiki/Lithornis

    It wasn't until 1988 that Houde, with a well preserved skull of *Lithornis* to study,
    found that it was palaeognathous. This put it very far from vultures, which
    are neognaths: there is a split right at the base of the tree of birds that survived the K-T disaster, with paleognaths on one side and neognaths
    on the other, according to most paleontologists specializing in birds,
    and most systematists, who use cladistic methods to study phylogeny.

    The palaeognaths of today comprise all the flightless birds known as ratites, and the tinamous, which are capable of short flights.

    The lithornids were also palaeognathous, but they were strong flyers
    capable of covering big distances without alighting. The ratites and
    the tinamous are both believed to be descended from them.

    They thus account for the widely separated places where ratites
    (ostrich, emu, cassowary, rhea, kiwi) are to be found. New Zealand
    was never connected to the continents where the other ratites
    are to be found. Its ratites included the moas along with the kiwis.


    Peter Nyikos
    Professor, Dept. of Mathematics -- standard disclaimer--
    Univ. of South Carolina in Columbia
    http://people.math.sc.edu/nyikos

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From erik simpson@21:1/5 to peter2...@gmail.com on Thu Nov 10 15:35:39 2022
    On Thursday, November 10, 2022 at 2:20:14 PM UTC-8, peter2...@gmail.com wrote:
    The lithornids, about which I wrote in the preceding post to this thread,
    got their name (literally, "stone birds") from the fact that the first one
    to be described, *Lithornis,* was among the very first fossil birds
    to be described in the scientific literature.

    Owen described it in 1840, long before anyone had heard of Archaeopteryx.
    He thought it was a vulture.

    https://en.wikipedia.org/wiki/Lithornis

    It wasn't until 1988 that Houde, with a well preserved skull of *Lithornis* to study,
    found that it was palaeognathous. This put it very far from vultures, which are neognaths: there is a split right at the base of the tree of birds that survived the K-T disaster, with paleognaths on one side and neognaths
    on the other, according to most paleontologists specializing in birds,
    and most systematists, who use cladistic methods to study phylogeny.

    The palaeognaths of today comprise all the flightless birds known as ratites, and the tinamous, which are capable of short flights.

    The lithornids were also palaeognathous, but they were strong flyers
    capable of covering big distances without alighting. The ratites and
    the tinamous are both believed to be descended from them.

    They thus account for the widely separated places where ratites
    (ostrich, emu, cassowary, rhea, kiwi) are to be found. New Zealand
    was never connected to the continents where the other ratites
    are to be found. Its ratites included the moas along with the kiwis.
    Peter Nyikos
    Professor, Dept. of Mathematics -- standard disclaimer--
    Univ. of South Carolina in Columbia
    http://people.math.sc.edu/nyikos

    You might find this interesting:

    https://digitallibrary.amnh.org/bitstream/handle/2246/7237/440-05-field_et_al.pdf?sequence=10&isAllowed=y

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From Trolidan7@21:1/5 to Peter Nyikos on Sun Nov 13 04:07:08 2022
    On 11/9/22 6:14 PM, Peter Nyikos wrote:
    The Great K-T extinction spelled the doom of the giant sea reptiles, all dinosaurs
    (unless you count birds as dinosaurs), and many other groups of vertebrates and invertebrates.
    In fact, the damage caused by the asteroid that hit Yucatan was so great
    that no animal over 50 kilos is known to have survived.

    But some groups of birds and mammals did survive, including the remote ancestors
    of the four groups that have survived to this day: the neornithine birds and three groups of mammals:
    the placentals, the marsupials, and the monotremes.

    To keep this OP reasonably short, I will deal with the two most familiar of the four, the placentals and the modern birds.

    A 2013 _Science_ article [1] led to the general acceptance of the hypothesis that all living placentals
    have descended from a common ancestor that lived after the great K-T disaster.

    However, there are closely related eutherians [2] that also survived for a while, among
    whose descendants were the pantodonts, including *Coryphodon*, which was a bit larger than a tapir; and the tillodonts and taeniodonts. These groups died out during the Eocene, which ended almost halfway between the K-T disaster and our time.

    The case of birds is generally believed to be quite different: EVERY bird, living
    or extinct, that survived the K-T disaster is believed to be descended from the
    last common ancestor of living birds. However, there are some articles which hypothesize that there is one exception, a group of birds called the lithornids.
    More about them in my next post to this thread, some time tomorrow.

    [1] Maureen A. O'Leary et. al., "The Placental Mammal Ancestor and the Post–K-Pg Radiation of Placentals". Science. 339 (6120): 662–667.

    [2] Eutherians are the mammals, extant and extinct, that are more closely related to placentals than to marsupials.
    Similarly, metatherians are the mammals that are more closely related to marsupials than to placentals.


    Peter Nyikos
    Professor, Dept. of Mathematics -- standard disclaimer--
    University of South Carolina
    http://people.math.sc.edu/nyikos

    PS The "Why?" at the end of the thread title is two-edged: why did some survive,
    and why did others, including some similar-seeming ones, perish?

    What sort of data did the 2013 Science article use to come to its
    conclusions? Was later widespread acceptance of the conclusions of the
    article based upon generally sound evidence?

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From erik simpson@21:1/5 to All on Sun Nov 13 10:14:41 2022
    On Sunday, November 13, 2022 at 4:07:11 AM UTC-8, Trolidan7 wrote:
    On 11/9/22 6:14 PM, Peter Nyikos wrote:
    The Great K-T extinction spelled the doom of the giant sea reptiles, all dinosaurs
    (unless you count birds as dinosaurs), and many other groups of vertebrates and invertebrates.
    In fact, the damage caused by the asteroid that hit Yucatan was so great that no animal over 50 kilos is known to have survived.

    But some groups of birds and mammals did survive, including the remote ancestors
    of the four groups that have survived to this day: the neornithine birds and three groups of mammals:
    the placentals, the marsupials, and the monotremes.

    To keep this OP reasonably short, I will deal with the two most familiar of the four, the placentals and the modern birds.

    A 2013 _Science_ article [1] led to the general acceptance of the hypothesis that all living placentals
    have descended from a common ancestor that lived after the great K-T disaster.

    However, there are closely related eutherians [2] that also survived for a while, among
    whose descendants were the pantodonts, including *Coryphodon*, which was a bit larger than a tapir; and the tillodonts and taeniodonts. These groups died out during the Eocene, which ended almost halfway between the K-T disaster and our time.

    The case of birds is generally believed to be quite different: EVERY bird, living
    or extinct, that survived the K-T disaster is believed to be descended from the
    last common ancestor of living birds. However, there are some articles which
    hypothesize that there is one exception, a group of birds called the lithornids.
    More about them in my next post to this thread, some time tomorrow.

    [1] Maureen A. O'Leary et. al., "The Placental Mammal Ancestor and the Post–K-Pg Radiation of Placentals". Science. 339 (6120): 662–667.

    [2] Eutherians are the mammals, extant and extinct, that are more closely related to placentals than to marsupials.
    Similarly, metatherians are the mammals that are more closely related to marsupials than to placentals.


    Peter Nyikos
    Professor, Dept. of Mathematics -- standard disclaimer--
    University of South Carolina
    http://people.math.sc.edu/nyikos

    PS The "Why?" at the end of the thread title is two-edged: why did some survive,
    and why did others, including some similar-seeming ones, perish?
    What sort of data did the 2013 Science article use to come to its conclusions? Was later widespread acceptance of the conclusions of the article based upon generally sound evidence?

    More recent research in placental evolution is reviewed here:

    "Evolutionary Models for the Diversification of Placental Mammals Across the KPg Boundary" (Springer et. al.)
    https://www.frontiersin.org/articles/10.3389/fgene.2019.01241/full

    The upshot is that "consensus on the tempo and mode of placental diversification remains elusive".

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From Trolidan7@21:1/5 to erik simpson on Mon Nov 14 11:13:45 2022
    On 11/13/22 10:14 AM, erik simpson wrote:
    On Sunday, November 13, 2022 at 4:07:11 AM UTC-8, Trolidan7 wrote:
    On 11/9/22 6:14 PM, Peter Nyikos wrote:
    The Great K-T extinction spelled the doom of the giant sea reptiles, all dinosaurs
    (unless you count birds as dinosaurs), and many other groups of vertebrates and invertebrates.
    In fact, the damage caused by the asteroid that hit Yucatan was so great >>> that no animal over 50 kilos is known to have survived.

    But some groups of birds and mammals did survive, including the remote
    ancestors
    of the four groups that have survived to this day: the neornithine birds and three groups of mammals:
    the placentals, the marsupials, and the monotremes.

    To keep this OP reasonably short, I will deal with the two most familiar of the four, the placentals and the modern birds.

    A 2013 _Science_ article [1] led to the general acceptance of the hypothesis that all living placentals
    have descended from a common ancestor that lived after the great K-T disaster.

    However, there are closely related eutherians [2] that also survived for a while, among
    whose descendants were the pantodonts, including *Coryphodon*, which was a bit larger than a tapir; and the tillodonts and taeniodonts. These groups died out during the Eocene, which ended almost halfway between the K-T disaster and our time.

    The case of birds is generally believed to be quite different: EVERY bird, living
    or extinct, that survived the K-T disaster is believed to be descended from the
    last common ancestor of living birds. However, there are some articles which
    hypothesize that there is one exception, a group of birds called the lithornids.
    More about them in my next post to this thread, some time tomorrow.

    [1] Maureen A. O'Leary et. al., "The Placental Mammal Ancestor and the Post–K-Pg Radiation of Placentals". Science. 339 (6120): 662–667.

    [2] Eutherians are the mammals, extant and extinct, that are more closely related to placentals than to marsupials.
    Similarly, metatherians are the mammals that are more closely related to marsupials than to placentals.


    Peter Nyikos
    Professor, Dept. of Mathematics -- standard disclaimer--
    University of South Carolina
    http://people.math.sc.edu/nyikos

    PS The "Why?" at the end of the thread title is two-edged: why did some survive,
    and why did others, including some similar-seeming ones, perish?
    What sort of data did the 2013 Science article use to come to its
    conclusions? Was later widespread acceptance of the conclusions of the
    article based upon generally sound evidence?

    More recent research in placental evolution is reviewed here:

    "Evolutionary Models for the Diversification of Placental Mammals Across the KPg Boundary" (Springer et. al.)
    https://www.frontiersin.org/articles/10.3389/fgene.2019.01241/full

    The upshot is that "consensus on the tempo and mode of placental diversification remains elusive".

    It is rather good that you can simply click on the link and get to
    the article.

    I have written down the Science 2013 article and see if I can
    find it somewhere.

    The articles that you have linked to seem to give the idea
    that 'we don't know'. That seems quite likely but I will
    see if I can find the Science article and see what it might
    say.

    I am thinking someone somewhere typed that 'penguins are
    the best of birds'. I would tend to disagree, for if
    there is something that could be called 'advanced' or
    'not advanced' I tend to like the song birds or passerines
    with their opposable toes for perching.

    When looking up the Wikipedia article on the song birds,
    however, it seemed to show some charts speculating that they
    might have diverged after the KPg boundary, but gave
    the idea that the ratites, land fowl, and water fowl
    may have all diverged prior to that. I would
    tend to think that if any of them diverged during the
    Mesozoic that it would be the Paleognaths.

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From erik simpson@21:1/5 to All on Mon Nov 14 13:25:46 2022
    On Monday, November 14, 2022 at 11:13:48 AM UTC-8, Trolidan7 wrote:
    On 11/13/22 10:14 AM, erik simpson wrote:
    On Sunday, November 13, 2022 at 4:07:11 AM UTC-8, Trolidan7 wrote:
    On 11/9/22 6:14 PM, Peter Nyikos wrote:
    The Great K-T extinction spelled the doom of the giant sea reptiles, all dinosaurs
    (unless you count birds as dinosaurs), and many other groups of vertebrates and invertebrates.
    In fact, the damage caused by the asteroid that hit Yucatan was so great >>> that no animal over 50 kilos is known to have survived.

    But some groups of birds and mammals did survive, including the remote >>> ancestors
    of the four groups that have survived to this day: the neornithine birds and three groups of mammals:
    the placentals, the marsupials, and the monotremes.

    To keep this OP reasonably short, I will deal with the two most familiar of the four, the placentals and the modern birds.

    A 2013 _Science_ article [1] led to the general acceptance of the hypothesis that all living placentals
    have descended from a common ancestor that lived after the great K-T disaster.

    However, there are closely related eutherians [2] that also survived for a while, among
    whose descendants were the pantodonts, including *Coryphodon*, which was a bit larger than a tapir; and the tillodonts and taeniodonts. These groups died out during the Eocene, which ended almost halfway between the K-T disaster and our time.

    The case of birds is generally believed to be quite different: EVERY bird, living
    or extinct, that survived the K-T disaster is believed to be descended from the
    last common ancestor of living birds. However, there are some articles which
    hypothesize that there is one exception, a group of birds called the lithornids.
    More about them in my next post to this thread, some time tomorrow.

    [1] Maureen A. O'Leary et. al., "The Placental Mammal Ancestor and the Post–K-Pg Radiation of Placentals". Science. 339 (6120): 662–667.

    [2] Eutherians are the mammals, extant and extinct, that are more closely related to placentals than to marsupials.
    Similarly, metatherians are the mammals that are more closely related to marsupials than to placentals.


    Peter Nyikos
    Professor, Dept. of Mathematics -- standard disclaimer--
    University of South Carolina
    http://people.math.sc.edu/nyikos

    PS The "Why?" at the end of the thread title is two-edged: why did some survive,
    and why did others, including some similar-seeming ones, perish?
    What sort of data did the 2013 Science article use to come to its
    conclusions? Was later widespread acceptance of the conclusions of the
    article based upon generally sound evidence?

    More recent research in placental evolution is reviewed here:

    "Evolutionary Models for the Diversification of Placental Mammals Across the KPg Boundary" (Springer et. al.)
    https://www.frontiersin.org/articles/10.3389/fgene.2019.01241/full

    The upshot is that "consensus on the tempo and mode of placental diversification remains elusive".
    It is rather good that you can simply click on the link and get to
    the article.

    I have written down the Science 2013 article and see if I can
    find it somewhere.

    The articles that you have linked to seem to give the idea
    that 'we don't know'. That seems quite likely but I will
    see if I can find the Science article and see what it might
    say.

    I am thinking someone somewhere typed that 'penguins are
    the best of birds'. I would tend to disagree, for if
    there is something that could be called 'advanced' or
    'not advanced' I tend to like the song birds or passerines
    with their opposable toes for perching.

    When looking up the Wikipedia article on the song birds,
    however, it seemed to show some charts speculating that they
    might have diverged after the KPg boundary, but gave
    the idea that the ratites, land fowl, and water fowl
    may have all diverged prior to that. I would
    tend to think that if any of them diverged during the
    Mesozoic that it would be the Paleognaths.

    Here's the O'Leary et.al. (2013):

    https://ri.conicet.gov.ar/bitstream/handle/11336/7302/CONICET_Digital_Nro.9756_A.pdf;jsessionid=10A2C6B4CF23B472582CCA7A281A2C14?sequence=2

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From Peter Nyikos@21:1/5 to All on Tue Nov 15 19:56:05 2022
    On Monday, November 14, 2022 at 2:13:48 PM UTC-5, Trolidan7 wrote:
    On 11/13/22 10:14 AM, erik simpson wrote:
    On Sunday, November 13, 2022 at 4:07:11 AM UTC-8, Trolidan7 wrote:
    On 11/9/22 6:14 PM, Peter Nyikos wrote:
    The Great K-T extinction spelled the doom of the giant sea reptiles, all dinosaurs
    (unless you count birds as dinosaurs), and many other groups of vertebrates and invertebrates.
    In fact, the damage caused by the asteroid that hit Yucatan was so great >>> that no animal over 50 kilos is known to have survived.

    But some groups of birds and mammals did survive, including the remote >>> ancestors
    of the four groups that have survived to this day: the neornithine birds and three groups of mammals:
    the placentals, the marsupials, and the monotremes.

    To keep this OP reasonably short, I will deal with the two most familiar of the four, the placentals and the modern birds.

    A 2013 _Science_ article [1] led to the general acceptance of the hypothesis that all living placentals
    have descended from a common ancestor that lived after the great K-T disaster.

    However, there are closely related eutherians [2] that also survived for a while, among
    whose descendants were the pantodonts, including *Coryphodon*, which was a bit larger than a tapir; and the tillodonts and taeniodonts. These groups died out during the Eocene, which ended almost halfway between the K-T disaster and our time.

    The case of birds is generally believed to be quite different: EVERY bird, living
    or extinct, that survived the K-T disaster is believed to be descended from the
    last common ancestor of living birds. However, there are some articles which
    hypothesize that there is one exception, a group of birds called the lithornids.
    More about them in my next post to this thread, some time tomorrow.

    [1] Maureen A. O'Leary et. al., "The Placental Mammal Ancestor and the Post–K-Pg Radiation of Placentals". Science. 339 (6120): 662–667.

    [2] Eutherians are the mammals, extant and extinct, that are more closely related to placentals than to marsupials.
    Similarly, metatherians are the mammals that are more closely related to marsupials than to placentals.


    Peter Nyikos
    Professor, Dept. of Mathematics -- standard disclaimer--
    University of South Carolina
    http://people.math.sc.edu/nyikos

    PS The "Why?" at the end of the thread title is two-edged: why did some survive,
    and why did others, including some similar-seeming ones, perish?
    What sort of data did the 2013 Science article use to come to its
    conclusions? Was later widespread acceptance of the conclusions of the
    article based upon generally sound evidence?

    More recent research in placental evolution is reviewed here:

    "Evolutionary Models for the Diversification of Placental Mammals Across the KPg Boundary" (Springer et. al.)
    https://www.frontiersin.org/articles/10.3389/fgene.2019.01241/full

    The upshot is that "consensus on the tempo and mode of placental diversification remains elusive".


    It is rather good that you can simply click on the link and get to
    the article.

    I have written down the Science 2013 article and see if I can
    find it somewhere.

    Erik gave you a link that works. I call your attention especially to the very detailed
    phylogenetic tree on page 663 (the second page of the article).

    That article has a colorful history.

    Almost immediately, some biologists, including at least one paleontologist, noted that Protungulatum, which appears on the tree just below the bats,
    also had another species that appeared during the late Cretaceous.

    This seemed to kill the central finding of the article, which claimed that
    the last common ancestor (LCA) of all living placentals appeared after
    the K-T boundary, and hence after the Cretaceous.

    However, a later and much more detailed phylogenetic analysis
    showed Protungulatum as not descended from that LCA.
    Unfortunately, I can't remember the title or the authors of
    that later research article. My printout of that article was packed away
    and I haven't found it yet. It did, however, appear before 2019,
    so it probably is covered somewhere in the Frontiers in Science
    article Erik linked for you [see above].

    The articles that you have linked to seem to give the idea
    that 'we don't know'. That seems quite likely but I will
    see if I can find the Science article and see what it might
    say.

    Unfortunately, as I indicated above, time has marched on
    in the world of research. The article Erik linked MIGHT
    have the last word in the line of argument I gave above,
    but the information in the article is so mashed together, I'm not sure
    I can find it, what with not remembering the title of that later article.


    Now you shifted from mammals to birds:

    I am thinking someone somewhere typed that 'penguins are
    the best of birds'.

    Whatever that means.

    I would tend to disagree, for if
    there is something that could be called 'advanced' or
    'not advanced' I tend to like the song birds or passerines
    with their opposable toes for perching.

    When looking up the Wikipedia article on the song birds,
    however, it seemed to show some charts speculating that they
    might have diverged after the KPg boundary, but gave
    the idea that the ratites, land fowl, and water fowl
    may have all diverged prior to that.

    Harshman is of this opinion.

    I would
    tend to think that if any of them diverged during the
    Mesozoic that it would be the Paleognaths.

    That does seem likely, but so far, no fossil paleognaths
    have turned up.


    Peter Nyikos
    Professor, Dept. of Mathematics
    Univ. of South Carolina -- standard disclaimer-- http://people.math.sc.edu/nyikos

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From erik simpson@21:1/5 to peter2...@gmail.com on Wed Nov 16 08:18:44 2022
    On Tuesday, November 15, 2022 at 7:56:06 PM UTC-8, peter2...@gmail.com wrote:
    ...
    Unfortunately, as I indicated above, time has marched on
    in the world of research.
    ...
    This isn't TO (fortunately), but this would otherwise be a great Chez Watt.

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From Peter Nyikos@21:1/5 to erik simpson on Wed Nov 16 09:48:36 2022
    On Wednesday, November 16, 2022 at 11:18:45 AM UTC-5, erik simpson wrote:
    On Tuesday, November 15, 2022 at 7:56:06 PM UTC-8, peter2...@gmail.com wrote:
    ...
    Unfortunately, as I indicated above, time has marched on
    in the world of research.
    ...
    This isn't TO (fortunately), but this would otherwise be a great Chez Watt.

    Not sure why: what category do you envision?

    Do keep in mind that a great Chez Watt does NOT rely on people knowing who uttered it.
    Nor, *a fortiori*, does it bank on reputations of the utterer based on unsupported canards in
    completely different contexts than the one specified with "as I indicated above."


    Anyway, I haven't had the chance to fully catch up with the march of time myself :)
    in that context, but I do have a lot of progress to report: I've found the article [1]
    that seems to have rehabilitated the central thesis of the 2013 Science article,
    that the crown group placentals had their LCA after the K-T disaster.

    [1] Halliday, Thomas John Dixon; Upchurch, Paul; Goswami, Anjali (2016-06-29). "Eutherians experienced elevated evolutionary rates in the immediate aftermath of the Cretaceous–Palaeogene mass extinction". Proceedings of the Royal Society B: Biological
    Sciences. 283 (1833): 20153026. https://discovery.ucl.ac.uk/id/eprint/1473028/1/Halliday_et_al-Biological_Reviews.pdf

    The article you linked [2] has no specific criticism for this paper,
    nor for a 2019 paper by Halliday et.al, [3] which supported the rehabilitation.
    However, it does have some very general cautionary notes,
    adding a more specific one from this same 2019 paper itself [4].

    [2] "Evolutionary Models for the Diversification of Placental Mammals Across the KPg Boundary" (Springer et. al.)
    https://www.frontiersin.org/articles/10.3389/fgene.2019.01241/full

    [3] Halliday, T. J., dos Reis, M., Tamuri, A. U., Ferguson-Gow, H., Yang, Z., Goswami, A. (2019). Rapid morphological evolution in placental mammals post-dates the origin of the crown group. Proc. R. Soc. Lond. B 286, 20182418. doi: 10.1098/rspb.2018.2418

    [4] "Finally, a recent study on morphological evolution in placental mammals concluded that it may be very difficult to distinguish early members of the major placental groups from stem eutherians on the basis of skeletal and dental characters because
    Cretaceous forms were not ecologically diverse and may appear very similar to each other (Halliday et al., 2019)."


    Temporary assessment: the post-K-T crown placental LCA hypothesis still seems to be the favored one, but nothing like a confident consensus has emerged.


    Peter Nyikos
    Professor, Dept. of Mathematics -- standard disclaimer--
    University of So. Carolina at Columbia
    http://people.math.sc.edu/nyikos

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From erik simpson@21:1/5 to peter2...@gmail.com on Wed Nov 16 10:17:34 2022
    On Wednesday, November 16, 2022 at 9:48:38 AM UTC-8, peter2...@gmail.com wrote:
    On Wednesday, November 16, 2022 at 11:18:45 AM UTC-5, erik simpson wrote:
    On Tuesday, November 15, 2022 at 7:56:06 PM UTC-8, peter2...@gmail.com wrote:
    ...
    Unfortunately, as I indicated above, time has marched on
    in the world of research.
    ...
    This isn't TO (fortunately), but this would otherwise be a great Chez Watt.
    Not sure why: what category do you envision?

    Do keep in mind that a great Chez Watt does NOT rely on people knowing who uttered it.
    Nor, *a fortiori*, does it bank on reputations of the utterer based on unsupported canards in
    completely different contexts than the one specified with "as I indicated above."


    Anyway, I haven't had the chance to fully catch up with the march of time myself :)
    in that context, but I do have a lot of progress to report: I've found the article [1]
    that seems to have rehabilitated the central thesis of the 2013 Science article,
    that the crown group placentals had their LCA after the K-T disaster.

    [1] Halliday, Thomas John Dixon; Upchurch, Paul; Goswami, Anjali (2016-06-29). "Eutherians experienced elevated evolutionary rates in the immediate aftermath of the Cretaceous–Palaeogene mass extinction". Proceedings of the Royal Society B:
    Biological Sciences. 283 (1833): 20153026.
    https://discovery.ucl.ac.uk/id/eprint/1473028/1/Halliday_et_al-Biological_Reviews.pdf

    The article you linked [2] has no specific criticism for this paper,
    nor for a 2019 paper by Halliday et.al, [3] which supported the rehabilitation.
    However, it does have some very general cautionary notes,
    adding a more specific one from this same 2019 paper itself [4].

    [2] "Evolutionary Models for the Diversification of Placental Mammals Across the KPg Boundary" (Springer et. al.)
    https://www.frontiersin.org/articles/10.3389/fgene.2019.01241/full

    [3] Halliday, T. J., dos Reis, M., Tamuri, A. U., Ferguson-Gow, H., Yang, Z., Goswami, A. (2019). Rapid morphological evolution in placental mammals post-dates the origin of the crown group. Proc. R. Soc. Lond. B 286, 20182418. doi: 10.1098/rspb.2018.
    2418

    [4] "Finally, a recent study on morphological evolution in placental mammals concluded that it may be very difficult to distinguish early members of the major placental groups from stem eutherians on the basis of skeletal and dental characters because
    Cretaceous forms were not ecologically diverse and may appear very similar to each other (Halliday et al., 2019)."


    Temporary assessment: the post-K-T crown placental LCA hypothesis still seems
    to be the favored one, but nothing like a confident consensus has emerged. Peter Nyikos
    Professor, Dept. of Mathematics -- standard disclaimer--
    University of So. Carolina at Columbia
    http://people.math.sc.edu/nyikos

    I don't believe you actually meant to say it's unfortunate that "time has marched on
    in the world of research", but you did say it. There's no direct connection to your reputation
    intended.

    Review papers are just that; summaries of recent work in some area of interest, primarily
    intended for those interested, but not intimately involved. Uncertainty is inevitable in periods
    of rapid evolution, such as follow major extinction events. Future discoveries may clarify some
    aspects, or may not. Fossils are sometimes reluctant witnesses.

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From Peter Nyikos@21:1/5 to erik simpson on Wed Nov 16 11:45:24 2022
    On Wednesday, November 16, 2022 at 1:17:35 PM UTC-5, erik simpson wrote:
    On Wednesday, November 16, 2022 at 9:48:38 AM UTC-8, peter2...@gmail.com wrote:
    On Wednesday, November 16, 2022 at 11:18:45 AM UTC-5, erik simpson wrote:
    On Tuesday, November 15, 2022 at 7:56:06 PM UTC-8, peter2...@gmail.com wrote:
    ...
    Unfortunately, as I indicated above, time has marched on
    in the world of research.
    ...
    This isn't TO (fortunately), but this would otherwise be a great Chez Watt.
    Not sure why: what category do you envision?

    Do keep in mind that a great Chez Watt does NOT rely on people knowing who uttered it.
    Nor, *a fortiori*, does it bank on reputations of the utterer based on unsupported canards in
    completely different contexts than the one specified with "as I indicated above."


    Anyway, I haven't had the chance to fully catch up with the march of time myself :)
    in that context, but I do have a lot of progress to report: I've found the article [1]
    that seems to have rehabilitated the central thesis of the 2013 Science article,
    that the crown group placentals had their LCA after the K-T disaster.

    [1] Halliday, Thomas John Dixon; Upchurch, Paul; Goswami, Anjali (2016-06-29). "Eutherians experienced elevated evolutionary rates in the immediate aftermath of the Cretaceous–Palaeogene mass extinction". Proceedings of the Royal Society B:
    Biological Sciences. 283 (1833): 20153026.
    https://discovery.ucl.ac.uk/id/eprint/1473028/1/Halliday_et_al-Biological_Reviews.pdf

    The article you linked [2] has no specific criticism for this paper,
    nor for a 2019 paper by Halliday et.al, [3] which supported the rehabilitation.
    However, it does have some very general cautionary notes,
    adding a more specific one from this same 2019 paper itself [4].

    [2] "Evolutionary Models for the Diversification of Placental Mammals Across the KPg Boundary" (Springer et. al.)
    https://www.frontiersin.org/articles/10.3389/fgene.2019.01241/full

    [3] Halliday, T. J., dos Reis, M., Tamuri, A. U., Ferguson-Gow, H., Yang, Z., Goswami, A. (2019). Rapid morphological evolution in placental mammals post-dates the origin of the crown group. Proc. R. Soc. Lond. B 286, 20182418. doi: 10.1098/rspb.2018.
    2418

    [4] "Finally, a recent study on morphological evolution in placental mammals concluded that it may be very difficult to distinguish early members of the major placental groups from stem eutherians on the basis of skeletal and dental characters
    because Cretaceous forms were not ecologically diverse and may appear very similar to each other (Halliday et al., 2019)."


    Temporary assessment: the post-K-T crown placental LCA hypothesis still seems
    to be the favored one, but nothing like a confident consensus has emerged. Peter Nyikos
    Professor, Dept. of Mathematics -- standard disclaimer--
    University of So. Carolina at Columbia
    http://people.math.sc.edu/nyikos

    I don't believe you actually meant to say it's unfortunate that "time has marched on
    in the world of research", but you did say it.

    You should believe it, because I only meant it in the context of what Trolidus7
    had said to you, which you snipped:

    "The articles that you have linked to seem to give the idea that 'we don't know'. That seems quite likely
    but I will see if I can find the Science article and see what it might say."

    The unfortunate part of the second sentence is that the 2013 Science article will not shed light
    on the first sentence, because one of the articles you linked came afterwards and was well aware of the pros and cons of the 2013 Science article.

    IOW, if I had not said what I did, Trolidus7 might have spent a lot of time before becoming
    disappointed over how little light the 2013 article shed on the issue of the first sentence.


    There's no direct connection to your reputation
    intended.

    That depends on how completely you forgot the part of the context that I recalled.


    Review papers are just that; summaries of recent work in some area of interest, primarily
    intended for those interested, but not intimately involved.

    Now that I've read it with keen attention to detail for the first time, that 2013 Science paper turns out to be
    just an extended research announcement, IOW a review paper on the authors' recent work
    that one hopes is sufficiently covered in the Supplementary materials.

    Unfortunately, :)
    the link you gave to that article does not give immediate direct link to the Supplementary Materials.
    Fortunately, I was able to get at them by the highlight-copy-and-paste-in-browser method:
    https://www.science.org/action/downloadSupplement?doi=10.1126%2Fscience.1229237&file=1229237oleary.sm.pdf

    This Supplement is much more detailed and complicated than the paper,
    and I doubt that I can get around to it until late next week.
    By then, John Harshman might be able to assess how good its use of various phylogenetic methods is.


    Uncertainty is inevitable in periods
    of rapid evolution, such as follow major extinction events.

    Not just rapid evolution, but rapid expansion in disparity. A big irony is that uncertainty seems
    to surround the most interesting and far-reaching expansions, such as the base of
    Theria, of Avialae, of Neornithes, of Archosauria, of Diapsida, of Synapsida, of Amniota,
    of Tetrapoda, of Vertebrata, and the most enigmatic of all: of Eumetazoa,
    all of which took place ca. the Cambrian Explosion.


    Future discoveries may clarify some
    aspects, or may not. Fossils are sometimes reluctant witnesses.

    All too true.


    Peter Nyikos
    Professor, Dept. of Mathematics -- standard disclaimer--
    University of So. Carolina in Columbia
    http://people.math.sc.edu/nyikos

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From Trolidan7@21:1/5 to Peter Nyikos on Wed Nov 16 16:05:52 2022
    On 11/15/22 7:56 PM, Peter Nyikos wrote:
    On Monday, November 14, 2022 at 2:13:48 PM UTC-5, Trolidan7 wrote:
    On 11/13/22 10:14 AM, erik simpson wrote:
    On Sunday, November 13, 2022 at 4:07:11 AM UTC-8, Trolidan7 wrote:
    On 11/9/22 6:14 PM, Peter Nyikos wrote:
    The Great K-T extinction spelled the doom of the giant sea reptiles, all dinosaurs
    (unless you count birds as dinosaurs), and many other groups of vertebrates and invertebrates.
    In fact, the damage caused by the asteroid that hit Yucatan was so great >>>>> that no animal over 50 kilos is known to have survived.

    But some groups of birds and mammals did survive, including the remote >>>>> ancestors
    of the four groups that have survived to this day: the neornithine birds and three groups of mammals:
    the placentals, the marsupials, and the monotremes.

    To keep this OP reasonably short, I will deal with the two most familiar of the four, the placentals and the modern birds.

    A 2013 _Science_ article [1] led to the general acceptance of the hypothesis that all living placentals
    have descended from a common ancestor that lived after the great K-T disaster.

    However, there are closely related eutherians [2] that also survived for a while, among
    whose descendants were the pantodonts, including *Coryphodon*, which was a bit larger than a tapir; and the tillodonts and taeniodonts. These groups died out during the Eocene, which ended almost halfway between the K-T disaster and our time.

    The case of birds is generally believed to be quite different: EVERY bird, living
    or extinct, that survived the K-T disaster is believed to be descended from the
    last common ancestor of living birds. However, there are some articles which
    hypothesize that there is one exception, a group of birds called the lithornids.
    More about them in my next post to this thread, some time tomorrow.

    [1] Maureen A. O'Leary et. al., "The Placental Mammal Ancestor and the Post–K-Pg Radiation of Placentals". Science. 339 (6120): 662–667.

    [2] Eutherians are the mammals, extant and extinct, that are more closely related to placentals than to marsupials.
    Similarly, metatherians are the mammals that are more closely related to marsupials than to placentals.


    Peter Nyikos
    Professor, Dept. of Mathematics -- standard disclaimer--
    University of South Carolina
    http://people.math.sc.edu/nyikos

    PS The "Why?" at the end of the thread title is two-edged: why did some survive,
    and why did others, including some similar-seeming ones, perish?
    What sort of data did the 2013 Science article use to come to its
    conclusions? Was later widespread acceptance of the conclusions of the >>>> article based upon generally sound evidence?

    More recent research in placental evolution is reviewed here:

    "Evolutionary Models for the Diversification of Placental Mammals Across the KPg Boundary" (Springer et. al.)
    https://www.frontiersin.org/articles/10.3389/fgene.2019.01241/full

    The upshot is that "consensus on the tempo and mode of placental diversification remains elusive".


    It is rather good that you can simply click on the link and get to
    the article.

    I have written down the Science 2013 article and see if I can
    find it somewhere.

    Erik gave you a link that works. I call your attention especially to the very detailed
    phylogenetic tree on page 663 (the second page of the article).

    That article has a colorful history.

    Almost immediately, some biologists, including at least one paleontologist, noted that Protungulatum, which appears on the tree just below the bats,
    also had another species that appeared during the late Cretaceous.

    This seemed to kill the central finding of the article, which claimed that the last common ancestor (LCA) of all living placentals appeared after
    the K-T boundary, and hence after the Cretaceous.

    However, a later and much more detailed phylogenetic analysis
    showed Protungulatum as not descended from that LCA.
    Unfortunately, I can't remember the title or the authors of
    that later research article. My printout of that article was packed away
    and I haven't found it yet. It did, however, appear before 2019,
    so it probably is covered somewhere in the Frontiers in Science
    article Erik linked for you [see above].

    The articles that you have linked to seem to give the idea
    that 'we don't know'. That seems quite likely but I will
    see if I can find the Science article and see what it might
    say.

    Unfortunately, as I indicated above, time has marched on
    in the world of research. The article Erik linked MIGHT
    have the last word in the line of argument I gave above,
    but the information in the article is so mashed together, I'm not sure
    I can find it, what with not remembering the title of that later article.


    Now you shifted from mammals to birds:

    I am thinking someone somewhere typed that 'penguins are
    the best of birds'.

    Whatever that means.

    I would tend to disagree, for if
    there is something that could be called 'advanced' or
    'not advanced' I tend to like the song birds or passerines
    with their opposable toes for perching.

    When looking up the Wikipedia article on the song birds,
    however, it seemed to show some charts speculating that they
    might have diverged after the KPg boundary, but gave
    the idea that the ratites, land fowl, and water fowl
    may have all diverged prior to that.

    Harshman is of this opinion.

    > I would
    tend to think that if any of them diverged during the
    Mesozoic that it would be the Paleognaths.

    That does seem likely, but so far, no fossil paleognaths
    have turned up.


    Peter Nyikos
    Professor, Dept. of Mathematics
    Univ. of South Carolina -- standard disclaimer-- http://people.math.sc.edu/nyikos

    It is amazing that you can come up with these articles
    that I can click on and see by just posting these links.

    At some point I am thinking I may want to tour some local
    libraries, however some time again.

    It would seem to me bad if it has been common in recent
    times for libraries to throw out such material thinking
    that internet forms are adequate, and then all the links
    mutated into something that could no longer be found or
    used because of paywalls or simply errors.

    Thank you both for the links and references.

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From Peter Nyikos@21:1/5 to All on Thu Nov 17 18:10:33 2022
    On Wednesday, November 16, 2022 at 7:05:55 PM UTC-5, Trolidan7 wrote:
    On 11/15/22 7:56 PM, Peter Nyikos wrote:
    On Monday, November 14, 2022 at 2:13:48 PM UTC-5, Trolidan7 wrote:
    On 11/13/22 10:14 AM, erik simpson wrote:
    On Sunday, November 13, 2022 at 4:07:11 AM UTC-8, Trolidan7 wrote:
    On 11/9/22 6:14 PM, Peter Nyikos wrote:
    The Great K-T extinction spelled the doom of the giant sea reptiles, all dinosaurs
    (unless you count birds as dinosaurs), and many other groups of vertebrates and invertebrates.
    In fact, the damage caused by the asteroid that hit Yucatan was so great
    that no animal over 50 kilos is known to have survived.

    But some groups of birds and mammals did survive, including the remote >>>>> ancestors
    of the four groups that have survived to this day: the neornithine birds and three groups of mammals:
    the placentals, the marsupials, and the monotremes.

    To keep this OP reasonably short, I will deal with the two most familiar of the four, the placentals and the modern birds.

    A 2013 _Science_ article [1] led to the general acceptance of the hypothesis that all living placentals
    have descended from a common ancestor that lived after the great K-T disaster.

    However, there are closely related eutherians [2] that also survived for a while, among
    whose descendants were the pantodonts, including *Coryphodon*, which was a bit larger than a tapir; and the tillodonts and taeniodonts. These groups died out during the Eocene, which ended almost halfway between the K-T disaster and our time.

    The case of birds is generally believed to be quite different: EVERY bird, living
    or extinct, that survived the K-T disaster is believed to be descended from the
    last common ancestor of living birds. However, there are some articles which
    hypothesize that there is one exception, a group of birds called the lithornids.
    More about them in my next post to this thread, some time tomorrow. >>>>>
    [1] Maureen A. O'Leary et. al., "The Placental Mammal Ancestor and the Post–K-Pg Radiation of Placentals". Science. 339 (6120): 662–667.

    [2] Eutherians are the mammals, extant and extinct, that are more closely related to placentals than to marsupials.
    Similarly, metatherians are the mammals that are more closely related to marsupials than to placentals.


    Peter Nyikos
    Professor, Dept. of Mathematics -- standard disclaimer--
    University of South Carolina
    http://people.math.sc.edu/nyikos

    PS The "Why?" at the end of the thread title is two-edged: why did some survive,
    and why did others, including some similar-seeming ones, perish?
    What sort of data did the 2013 Science article use to come to its
    conclusions? Was later widespread acceptance of the conclusions of the >>>> article based upon generally sound evidence?

    More recent research in placental evolution is reviewed here:

    "Evolutionary Models for the Diversification of Placental Mammals Across the KPg Boundary" (Springer et. al.)
    https://www.frontiersin.org/articles/10.3389/fgene.2019.01241/full

    The upshot is that "consensus on the tempo and mode of placental diversification remains elusive".


    It is rather good that you can simply click on the link and get to
    the article.

    I have written down the Science 2013 article and see if I can
    find it somewhere.

    Erik gave you a link that works. I call your attention especially to the very detailed
    phylogenetic tree on page 663 (the second page of the article).

    That article has a colorful history.

    Almost immediately, some biologists, including at least one paleontologist,
    noted that Protungulatum, which appears on the tree just below the bats, also had another species that appeared during the late Cretaceous.

    This seemed to kill the central finding of the article, which claimed that the last common ancestor (LCA) of all living placentals appeared after
    the K-T boundary, and hence after the Cretaceous.

    However, a later and much more detailed phylogenetic analysis
    showed Protungulatum as not descended from that LCA.
    Unfortunately, I can't remember the title or the authors of
    that later research article. My printout of that article was packed away and I haven't found it yet. It did, however, appear before 2019,
    so it probably is covered somewhere in the Frontiers in Science
    article Erik linked for you [see above].

    Happy update: Google was my friend: I found the article by looking up Protungulatum
    in Wikipedia and recognizing the name of the author in one of the references. It was a 2016 article by Halliday et. al. I gave full bibliographic information on it in my last
    reply I did to Erik yesterday, including a link to a fine digital copy.


    <snip for focus>


    When looking up the Wikipedia article on the song birds,
    however, it seemed to show some charts speculating that they
    might have diverged after the KPg boundary, but gave
    the idea that the ratites, land fowl, and water fowl
    may have all diverged prior to that.

    Harshman is of this opinion.

    I would
    tend to think that if any of them diverged during the
    Mesozoic that it would be the Paleognaths.

    That does seem likely, but so far, no fossil paleognaths
    have turned up.

    I meant, of course, no fossil paleognaths of the Cretaceous.


    It is amazing that you can come up with these articles
    that I can click on and see by just posting these links.

    There is sometimes some hard work in the background to
    find webpages where a copy of an article is not paywalled.
    I couldn't find one for the Science 2013 article, but Erik came
    through with one.

    There is also a matter of quality. At one time, I had access to
    a copy of that same article where that phylogenetic tree could be
    magnified. The one Erik found does not have that capability and
    so I have to sometimes use a magnifying glass to make out certain features.


    At some point I am thinking I may want to tour some local
    libraries, however some time again.

    It would seem to me bad if it has been common in recent
    times for libraries to throw out such material thinking
    that internet forms are adequate, and then all the links
    mutated into something that could no longer be found or
    used because of paywalls or simply errors.

    Libraries have been deteriorating in various ways. One small branch
    of the biggest library in the Columbia area has stopped using
    the Dewey decimal system and arranges books according to
    the names of authors! I hope that doesn't catch on elsewhere!

    Our university library has long had a habit of carting off books
    to a faraway annex if they have not been checked out in ten years.
    I used to go to the library at least once a year to look things up in
    a popular book in topology, but didn't see any reason to check it out.
    Then one day it was no longer available except by putting in a request to the annex.


    Thank you both for the links and references.

    Glad to be of help. I hope you keep following this thread,
    because I have lots more to contribute to it.
    Tomorrow, I will have a post on monotremes. Next week,
    Monday, I'll have another on birds; then come marsupials.


    Peter Nyikos

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From Popping Mad@21:1/5 to Peter Nyikos on Fri Nov 18 11:15:49 2022
    On 11/9/22 21:14, Peter Nyikos wrote:
    To keep this OP reasonably short, I will deal with the two most familiar of the four, the placentals and the modern birds.


    short?

    Frogs hopped right through.

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From erik simpson@21:1/5 to Popping Mad on Sun Nov 20 08:36:26 2022
    On Sunday, November 20, 2022 at 8:26:25 AM UTC-8, Popping Mad wrote:
    On 11/10/22 17:20, Peter Nyikos wrote:
    They thus account for the widely separated places where ratites
    (ostrich, emu, cassowary, rhea, kiwi) are to be found. New Zealand
    was never connected to the continents where the other ratites
    are to be found. Its ratites included the moas along with the kiwis.
    The real question is how would they taste on Thanksgiving!


    Good week one and all.

    Emus aren't bad, but they don't taste like turkey.

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From Popping Mad@21:1/5 to Peter Nyikos on Sun Nov 20 11:25:47 2022
    On 11/10/22 17:20, Peter Nyikos wrote:
    They thus account for the widely separated places where ratites
    (ostrich, emu, cassowary, rhea, kiwi) are to be found. New Zealand
    was never connected to the continents where the other ratites
    are to be found. Its ratites included the moas along with the kiwis.


    The real question is how would they taste on Thanksgiving!


    Good week one and all.

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From Popping Mad@21:1/5 to erik simpson on Sun Nov 20 18:48:37 2022
    On 11/20/22 11:36, erik simpson wrote:
    Emus aren't bad, but they don't taste like turkey.


    try roasted yellow canary in a puffery roll with a dab of powered sugar.
    It does great with Irish Coffee at the fireplace before the main course
    ...

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From erik simpson@21:1/5 to Popping Mad on Sun Nov 20 16:49:12 2022
    On Sunday, November 20, 2022 at 3:49:15 PM UTC-8, Popping Mad wrote:
    On 11/20/22 11:36, erik simpson wrote:
    Emus aren't bad, but they don't taste like turkey.
    try roasted yellow canary in a puffery roll with a dab of powered sugar.
    It does great with Irish Coffee at the fireplace before the main course
    ...
    Here's hoping you aren't serious! Emu is actully sold in groceries and listed on
    the menu of many restaurants in Oz. It's red meat, and as I said, not at all like
    turkey. Ostrich meat is available in the US, but I've never eaten any. I understand
    it's like emu; red, more similar in taste to beef than the birds we're more used to.

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From Peter Nyikos@21:1/5 to Popping Mad on Mon Nov 21 14:34:41 2022
    On Friday, November 18, 2022 at 11:16:28 AM UTC-5, Popping Mad wrote:
    On 11/9/22 21:14, Peter Nyikos wrote:
    To keep this OP reasonably short, I will deal with the two most familiar of the four, the placentals and the modern birds.
    short?

    Frogs hopped right through.

    I'm not familiar with that figure of speech.

    By the way, I started a thread last month whose title ended ATTN: Popping mad. The following link takes you to why I wanted to catch your attention:

    https://groups.google.com/g/sci.bio.paleontology/c/U-99grFea8E/m/fQjzHZNyAwAJ Re: Hesperornid Acquisition here in Columbia ATTN: Popping mad
    Oct 14, 2022, 7:54:33 PM

    Hope to see you over there.


    Peter Nyikos

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From Peter Nyikos@21:1/5 to Peter Nyikos on Wed Nov 23 19:26:05 2022
    On Thursday, November 17, 2022 at 9:10:34 PM UTC-5, Peter Nyikos wrote:
    On Wednesday, November 16, 2022 at 7:05:55 PM UTC-5, Trolidan7 wrote:

    Thank you both for the links and references.
    Glad to be of help. I hope you keep following this thread,
    because I have lots more to contribute to it.
    Tomorrow, I will have a post on monotremes. Next week,
    Monday, I'll have another on birds; then come marsupials.

    The best laid plans of mice and men...

    I hope you haven't given up waiting. Here is the post on monotremes; then tomorrow,
    unless my family has a very full schedule, comes the next one on birds.

    Monotremes have a very sparse fossil record, but it goes back to the early Cretaceous,
    over 100 mya, when there were at least three different genera: Steropodon, Kryoryctes, and Teinolophos.
    Then, except for some disputed specimens, there is nothing until the only non-Australian
    known monotreme, Monotrematum sudamericanum – 61 million years old, past the K-T
    extinction event. Some classify it as a species of Obdurodon, otherwise known from
    the early Miocene (24 mya) to the very late Miocene, 5 mya. If the identification
    is correct, then Obdurodon was a "living fossil" by the time it became extinct.

    Obdurodon was quite similar to the living platypus, except that it had some teeth
    as an adult. The living platypus, Ornithorhynchus anatinus, only has milk teeth and when those are shed, only the familiar bill remains.

    If Monotrematum was actually an Obdurodon, then it is quite probable
    that Obdurodon was a K-T survivor, the lone monotreme with that distinction.

    A funny thing about all these monotremes: they all are believed
    to be "duckbills" of some sort. The echidnas diverged from them
    at some time and have obviously undergone some drastic changes in their mouth parts.


    Peter Nyikos
    Professor, Dept. of Mathematics
    University of So. Carolina -- standard disclaimer-- http://people.math.sc.edu/nyikos

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From Trolidan7@21:1/5 to Peter Nyikos on Thu Nov 24 11:47:30 2022
    On 11/23/22 7:26 PM, Peter Nyikos wrote:
    On Thursday, November 17, 2022 at 9:10:34 PM UTC-5, Peter Nyikos wrote:
    On Wednesday, November 16, 2022 at 7:05:55 PM UTC-5, Trolidan7 wrote:

    Thank you both for the links and references.
    Glad to be of help. I hope you keep following this thread,
    because I have lots more to contribute to it.
    Tomorrow, I will have a post on monotremes. Next week,
    Monday, I'll have another on birds; then come marsupials.

    The best laid plans of mice and men...

    I hope you haven't given up waiting. Here is the post on monotremes; then tomorrow,
    unless my family has a very full schedule, comes the next one on birds.

    Monotremes have a very sparse fossil record, but it goes back to the early Cretaceous,
    over 100 mya, when there were at least three different genera: Steropodon, Kryoryctes, and Teinolophos.
    Then, except for some disputed specimens, there is nothing until the only non-Australian
    known monotreme, Monotrematum sudamericanum – 61 million years old, past the K-T
    extinction event. Some classify it as a species of Obdurodon, otherwise known from
    the early Miocene (24 mya) to the very late Miocene, 5 mya. If the identification
    is correct, then Obdurodon was a "living fossil" by the time it became extinct.

    Obdurodon was quite similar to the living platypus, except that it had some teeth
    as an adult. The living platypus, Ornithorhynchus anatinus, only has milk teeth
    and when those are shed, only the familiar bill remains.

    If Monotrematum was actually an Obdurodon, then it is quite probable
    that Obdurodon was a K-T survivor, the lone monotreme with that distinction.

    A funny thing about all these monotremes: they all are believed
    to be "duckbills" of some sort. The echidnas diverged from them
    at some time and have obviously undergone some drastic changes in their mouth parts.

    Well, you know, surfing it some gives me the idea that for a
    lot of this phenomena a lot of information is derived from
    fossil tooth structure. I get the idea that Multituberculata
    survived the K-T boundary, but they died out before the present.

    It seems feasible to me that if they had survived to the
    present they could have been classified as separate and
    distinct among the living mammals like the monotremes.

    Or would their reproductive strategies have been similar
    enough to either the monotremes, marsupials, or placental
    mammals for them to have been classified as curious members
    of one of the others until recently through genetics?

    I guess you can infer the existence of 'milk teeth'
    from fossils. It is reasonable that theoretically
    inferring from the existence of 'milk teeth' that members
    of multituberculata gave or drank milk at some point in
    their life cycle?

    Peter Nyikos
    Professor, Dept. of Mathematics
    University of So. Carolina -- standard disclaimer-- http://people.math.sc.edu/nyikos

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From Trolidan7@21:1/5 to All on Thu Nov 24 15:57:51 2022
    On 11/24/22 11:47 AM, Trolidan7 wrote:
    On 11/23/22 7:26 PM, Peter Nyikos wrote:
    On Thursday, November 17, 2022 at 9:10:34 PM UTC-5, Peter Nyikos wrote:
    On Wednesday, November 16, 2022 at 7:05:55 PM UTC-5, Trolidan7 wrote:

    Thank you both for the links and references.
    Glad to be of help. I hope you keep following this thread,
    because I have lots more to contribute to it.
    Tomorrow, I will have a post on monotremes. Next week,
    Monday, I'll have another on birds; then come marsupials.

    The best laid plans of mice and men...

    I hope you haven't given up waiting. Here is the post on monotremes;
    then tomorrow,
    unless my family has a very full schedule, comes the next one on birds.

    Monotremes have a very sparse fossil record, but it goes back to the
    early Cretaceous,
    over 100 mya, when there were at least three different genera:
    Steropodon, Kryoryctes, and Teinolophos.
    Then, except for some disputed specimens, there is nothing until the
    only non-Australian
    known monotreme, Monotrematum sudamericanum – 61 million years old,
    past the K-T
    extinction event. Some classify it as a species of Obdurodon,
    otherwise known from
    the early Miocene (24 mya) to the very late Miocene, 5 mya. If the
    identification
    is correct, then Obdurodon was a "living fossil" by the time it became
    extinct.

    Obdurodon was quite similar to the living platypus, except that it had
    some teeth
    as an adult. The living platypus, Ornithorhynchus anatinus, only has
    milk teeth
    and when those are shed, only the familiar bill remains.

    If Monotrematum was actually an Obdurodon, then it is quite probable
    that Obdurodon was a K-T survivor, the lone monotreme with that
    distinction.

    A funny thing about all these monotremes: they all are believed
    to be "duckbills" of some sort. The echidnas diverged from them
    at some time and have obviously undergone some drastic changes in
    their mouth parts.

    Well, you know, surfing it some gives me the idea that for a
    lot of this phenomena a lot of information is derived from
    fossil tooth structure.  I get the idea that Multituberculata
    survived the K-T boundary, but they died out before the present.

    It seems feasible to me that if they had survived to the
    present they could have been classified as separate and
    distinct among the living mammals like the monotremes.

    Or would their reproductive strategies have been similar
    enough to either the monotremes, marsupials, or placental
    mammals for them to have been classified as curious members
    of one of the others until recently through genetics?

    I guess you can infer the existence of 'milk teeth'
    from fossils.  It is reasonable that theoretically
    inferring from the existence of 'milk teeth' that members
    of multituberculata gave or drank milk at some point in
    their life cycle?

    You know, when I think about it, 'milk
    teeth' is just a name. It is possible
    for a person to consume a glass of milk
    without having any milk teeth.

    Who knows what these animals may have
    been like.

    Peter Nyikos
    Professor, Dept. of Mathematics
    University of So. Carolina  -- standard disclaimer--
    http://people.math.sc.edu/nyikos


    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From Peter Nyikos@21:1/5 to All on Thu Nov 24 17:46:38 2022
    On Thursday, November 24, 2022 at 2:47:34 PM UTC-5, Trolidan7 wrote:
    On 11/23/22 7:26 PM, Peter Nyikos wrote:
    On Thursday, November 17, 2022 at 9:10:34 PM UTC-5, Peter Nyikos wrote:
    On Wednesday, November 16, 2022 at 7:05:55 PM UTC-5, Trolidan7 wrote:

    Thank you both for the links and references.
    Glad to be of help. I hope you keep following this thread,
    because I have lots more to contribute to it.
    Tomorrow, I will have a post on monotremes. Next week,
    Monday, I'll have another on birds; then come marsupials.

    The best laid plans of mice and men...

    I hope you haven't given up waiting. Here is the post on monotremes; then tomorrow,
    unless my family has a very full schedule, comes the next one on birds.

    Monotremes have a very sparse fossil record, but it goes back to the early Cretaceous,
    over 100 mya, when there were at least three different genera: Steropodon, Kryoryctes, and Teinolophos.
    Then, except for some disputed specimens, there is nothing until the only non-Australian
    known monotreme, Monotrematum sudamericanum – 61 million years old, past the K-T
    extinction event. Some classify it as a species of Obdurodon, otherwise known from
    the early Miocene (24 mya) to the very late Miocene, 5 mya. If the identification
    is correct, then Obdurodon was a "living fossil" by the time it became extinct.

    Obdurodon was quite similar to the living platypus, except that it had some teeth
    as an adult. The living platypus, Ornithorhynchus anatinus, only has milk teeth
    and when those are shed, only the familiar bill remains.

    If Monotrematum was actually an Obdurodon, then it is quite probable
    that Obdurodon was a K-T survivor, the lone monotreme with that distinction.

    A funny thing about all these monotremes: they all are believed
    to be "duckbills" of some sort. The echidnas diverged from them
    at some time and have obviously undergone some drastic changes in their mouth parts.

    One strange feature of the Science 2013 article that Erik linked
    for us is that it puts the fork between platypuses and echidnas
    right about the time of the first confirmed fossil of *Obdurodon*,
    for no reason I could see. Since the first known fossils of
    echidnas came later, it puts in a light streak for "ghost taxa"
    between them and the fork.

    Unless the authors have some molecular clock arguments that I missed,
    they could just as easily have put the fork further back
    and had ghost taxa stretching from it to both families.

    Molecular clock arguments are a two-edged sword: some
    would invalidate the thesis that the placental crown group
    goes back to AFTER the K-T disaster, which the 2013 article endorses.


    Well, you know, surfing it some gives me the idea that for a
    lot of this phenomena a lot of information is derived from
    fossil tooth structure. I get the idea that Multituberculata
    survived the K-T boundary, but they died out before the present.

    Multituberculata is part of a separate infraclass of Mammalia from both Theria [1]
    and Prototheria (including Monotremata), and I was only going to look at it closely later. However, it is clear that several lines from it made it
    past the K-T boundary. In fact, the wiki entry says that they reached
    their greatest diversity in the Upper Cretaceous and Paleocene,
    almost as though the K-T disaster had hardly touched them.

    https://en.wikipedia.org/wiki/Multituberculata

    However, I intend to look deeper into this before
    making any definite claims.

    [1] placentals and marsupials (or eutherians and metatherians if
    you take into account extinct lineages after placentals split from marsupials).


    It seems feasible to me that if they had survived to the
    present they could have been classified as separate and
    distinct among the living mammals like the monotremes.

    Absolutely. This has been the general opinion of paleontologists
    for well over a century now, and the wiki entry goes into some of
    the differences.

    Or would their reproductive strategies have been similar
    enough to either the monotremes, marsupials, or placental
    mammals for them to have been classified as curious members
    of one of the others until recently through genetics?

    Reproductive strategies are surprisingly different even within
    Placentalia, and are not a good guide for relationships.

    It is even conceivable that monotremes are secondarily
    egg layers with a common ancestor that gave live birth.
    Their eggs seem to be unique in that they absorb complex
    nutrients from the uterus, and not just water and gases
    like with the eggs of almost all oviparous vertebrates.

    I guess you can infer the existence of 'milk teeth'
    from fossils.

    Yes, there is a big difference between the primary ("milk")
    teeth of mammals, which are shed at some point in development,
    and the permanent adult teeth. I believe any comparative
    anatomist can tell the difference.

    It is reasonable that theoretically
    inferring from the existence of 'milk teeth' that members
    of multituberculata gave or drank milk at some point in
    their life cycle?

    Not from teeth: mammals are *defined* among living vertebrates as
    giving milk from glands in the skin. The dividing line
    has to do with the bones in the ear, whether there is
    only the stapes as in reptiles (more relevantly, therapsids)
    or there are also the malleus and incus. These were
    formerly tiny jaw bones known as the angular and quadrate,
    which formed the therapsid jaw joint, and then migrated to the middle ear.


    Peter Nyikos
    Professor, Dept. of Mathematics -- standard disclaimer--
    University of So. Carolina at Columbia
    http://people.math.sc.edu/nyikos

    PS I forgot that my family's celebrating of Thanksgiving
    would cut into my posting time. I'll try to get back to
    the question of which bird lineages survived the K-T disaster
    tomorrow.

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From Peter Nyikos@21:1/5 to John Harshman on Fri Nov 25 19:56:25 2022
    This is my second reply to a post by John Harshman on another thread:

    https://groups.google.com/g/sci.bio.paleontology/c/U-99grFea8E/m/vj8TF0RaAAAJ Re: Hesperornid Acquisition here in Columbia ATTN: Popping mad

    That title long outlived its usefulness, and my first reply already
    would already have been more appropriate here. It was: https://groups.google.com/g/sci.bio.paleontology/c/U-99grFea8E/m/K0OXzSgIAQAJ


    On Wednesday, October 26, 2022 at 4:30:33 PM UTC-4, John Harshman wrote:
    On 10/26/22 12:57 PM, Peter Nyikos wrote:
    On Friday, October 21, 2022 at 10:14:35 PM UTC-4, John Harshman wrote:
    On 10/21/22 1:36 PM, Peter Nyikos wrote:
    On Friday, October 21, 2022 at 3:24:55 PM UTC-4, John Harshman wrote: >>>> On 10/26/22 12:57 PM, Peter Nyikos wrote:

    Picking up almost where I left off in my first reply:

    there are at least three bird lineages
    surviving the K/P extinction.

    Again, at least 7: paleognaths, neoaves, galliforms, anhimids,
    presbyornithids, anseranatidae, anatidae.

    Which bird did you have in mind from neoaves?

    It's worth noting that the last 5 groups you list are all in Galloanserae.
    If, as is generally believed, the first split in Neornithes is between paleognaths and Galloanserae, you are assuming below that
    Neoaves and Galloanserae are sister groups, right?


    It's all implied by the tree if we suppose there are Cretaceous
    presbyornithids and that presbyornithids are the sister group of
    anatids. I wasn't referring to any other fossils.

    The problem with that, as we saw in the first reply, is
    that the claim of there being Cretaceous presbyornithids
    is shaky. You need several generally accepted Cretaceous
    members of Galloanserae to make the claim that there
    were 7 lineages.


    This double-dipping (the same fossils for neoaves and presbyornithids) brings the total down to at most 6 but probably at most 5.

    Beg pardon? I fail to understand your objection here. The point is that
    if group X exists, so must its sister group.

    Right, but how far does this generality take us? We need to be clear on where we are relying on this group X - sister talk, and where we
    are relying on Cretaceous fossils.


    By the way, are you using Cretaceous representatives of crown groups?
    I mean crown anatiforms for the Cretaceous anatiforms, etc. (except of course
    for the presbyornithids, which became extinct in the Miocene).

    You mean "anseriforms" or possibly "anatids". Presbyornithids are crown
    group anseriforms. If there are Cretaceous presbyornithinds, then there
    must also be Cretaceous anatids, since they are sister groups.

    There is that nasty word "if" again. Where are the generally accepted Cretaceous fossils?


    Peter Nyikos
    Professor, Dept. of Mathematics -- standard disclaimer--
    Univ. of So. Carolina at Columbia
    http://people.math.sc.edu/nyikos

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From John Harshman@21:1/5 to Peter Nyikos on Sat Nov 26 22:07:04 2022
    On 11/25/22 7:56 PM, Peter Nyikos wrote:
    This is my second reply to a post by John Harshman on another thread:

    https://groups.google.com/g/sci.bio.paleontology/c/U-99grFea8E/m/vj8TF0RaAAAJ Re: Hesperornid Acquisition here in Columbia ATTN: Popping mad

    That title long outlived its usefulness, and my first reply already
    would already have been more appropriate here. It was: https://groups.google.com/g/sci.bio.paleontology/c/U-99grFea8E/m/K0OXzSgIAQAJ


    On Wednesday, October 26, 2022 at 4:30:33 PM UTC-4, John Harshman wrote:
    On 10/26/22 12:57 PM, Peter Nyikos wrote:
    On Friday, October 21, 2022 at 10:14:35 PM UTC-4, John Harshman wrote:
    On 10/21/22 1:36 PM, Peter Nyikos wrote:
    On Friday, October 21, 2022 at 3:24:55 PM UTC-4, John Harshman wrote: >>>>>> On 10/26/22 12:57 PM, Peter Nyikos wrote:

    Picking up almost where I left off in my first reply:

    there are at least three bird lineages
    surviving the K/P extinction.

    Again, at least 7: paleognaths, neoaves, galliforms, anhimids,
    presbyornithids, anseranatidae, anatidae.

    Which bird did you have in mind from neoaves?

    It's worth noting that the last 5 groups you list are all in Galloanserae. If, as is generally believed, the first split in Neornithes is between paleognaths and Galloanserae, you are assuming below that
    Neoaves and Galloanserae are sister groups, right?


    It's all implied by the tree if we suppose there are Cretaceous
    presbyornithids and that presbyornithids are the sister group of
    anatids. I wasn't referring to any other fossils.

    The problem with that, as we saw in the first reply, is
    that the claim of there being Cretaceous presbyornithids
    is shaky. You need several generally accepted Cretaceous
    members of Galloanserae to make the claim that there
    were 7 lineages.

    No, you only need one, as long as it's the right one. Either a
    presbyornithid or an anatid would do.

    Here is a claim of a Cretaceous presbyornithid:

    https://www.ncbi.nlm.nih.gov/pmc/articles/PMC4785986/

    This double-dipping (the same fossils for neoaves and presbyornithids)
    brings the total down to at most 6 but probably at most 5.

    Beg pardon? I fail to understand your objection here. The point is that
    if group X exists, so must its sister group.

    Right, but how far does this generality take us? We need to be clear on where we are relying on this group X - sister talk, and where we
    are relying on Cretaceous fossils.

    Again, one Cretaceous presbyornithid implies at least 7 lineages; Presbyornithidae, Anatidae, Anseranatidae, Anhimidae, Galliformes,
    Neoaves, Palaeognathae.

    One might also add Gastornithidae and Pelagornithidae, though I'm not
    sure whether those are both separate branches.

    By the way, are you using Cretaceous representatives of crown groups?
    I mean crown anatiforms for the Cretaceous anatiforms, etc. (except of course
    for the presbyornithids, which became extinct in the Miocene).

    You mean "anseriforms" or possibly "anatids". Presbyornithids are crown
    group anseriforms. If there are Cretaceous presbyornithinds, then there
    must also be Cretaceous anatids, since they are sister groups.

    There is that nasty word "if" again. Where are the generally accepted Cretaceous fossils?

    I'm not sure the literature on this is extensive enough for there to be
    any way to evaluate "generally accepted". One can only present publications.

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From Peter Nyikos@21:1/5 to John Harshman on Mon Nov 28 19:13:10 2022
    On Sunday, November 27, 2022 at 1:07:11 AM UTC-5, John Harshman wrote:
    On 11/25/22 7:56 PM, Peter Nyikos wrote:
    On Wednesday, October 26, 2022 at 4:30:33 PM UTC-4, John Harshman wrote:
    On 10/26/22 12:57 PM, Peter Nyikos wrote:
    On Friday, October 21, 2022 at 10:14:35 PM UTC-4, John Harshman wrote: >>>> On 10/21/22 1:36 PM, Peter Nyikos wrote:
    On Friday, October 21, 2022 at 3:24:55 PM UTC-4, John Harshman wrote: >>>>>> On 10/26/22 12:57 PM, Peter Nyikos wrote:

    there are at least three bird lineages
    surviving the K/P extinction.

    Again, at least 7: paleognaths, neoaves, galliforms, anhimids,
    presbyornithids, anseranatidae, anatidae.

    Which bird did you have in mind from neoaves?

    It's worth noting that the last 5 groups you list are all in Galloanserae.

    And the last four are generally believed to be in Anseriformes ("waterfowl").

    If, as is generally believed, the first split in Neornithes is between paleognaths and Galloanserae, you are assuming below that
    Neoaves and Galloanserae are sister groups, right?

    You don't answer this, but I soon figured out that I was right here.


    It's all implied by the tree if we suppose there are Cretaceous
    presbyornithids and that presbyornithids are the sister group of
    anatids. I wasn't referring to any other fossils.

    The problem with that, as we saw in the first reply, is
    that the claim of there being Cretaceous presbyornithids
    is shaky. You need several generally accepted Cretaceous
    members of Galloanserae to make the claim that there
    were 7 lineages.

    No, you only need one, as long as it's the right one. Either a presbyornithid or an anatid would do.

    You are assuming some other features of the phylogenetic tree
    of the order Anseriformes ("waterfowl") that you aren't
    identifying here. What are they?

    Here is a claim of a Cretaceous presbyornithid:

    https://www.ncbi.nlm.nih.gov/pmc/articles/PMC4785986/

    It sounds rather tentative, so not too encouraging of "only need one":

    "Although screamers (Anhimidae) may have evolved from presbyornithid-like birds, the uncertainty as to whether skeletal features shared by presbyornithids and anhimids are plesiomorphic within Anseriformes or indicative of a close relationship between
    the two cannot yet be resolved. In any case, recognition of W. tedfordi as a presbyornithid calls for a reassessment of the phylogenetic position of Presbyornithidae within Anseriformes."

    Sounds like your hypothesis is much more easily falsifiable than the hypothesis that a
    species of *Merychippus* is directly ancestral to *Equus* in the horse family.


    One might also add Gastornithidae and Pelagornithidae, though I'm not
    sure whether those are both separate branches.

    Ah, one of my favorite bird families, those pelagornithids ["pseudotoothed birds"].
    The biggest one of all, with a wingspread over 20 feet, was found right here
    in South Carolina, while clearing land for the Charleston international airport.


    However, I saw no candidates for either family having Cretaceous members.
    And whether they are separate branches or the same seems irrelevant.
    You really have your work cut out for you to find an indirect phylogenetic
    tree way to make them survivors of the K-P extinction.


    Peter Nyikos
    Professor, Dept. of Mathematics -- standard disclaimer--
    Univ. of So. Carolina at Columbia
    http://people.math.sc.edu/nyikos

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From John Harshman@21:1/5 to Peter Nyikos on Tue Nov 29 06:19:07 2022
    On 11/28/22 7:13 PM, Peter Nyikos wrote:
    On Sunday, November 27, 2022 at 1:07:11 AM UTC-5, John Harshman wrote:
    On 11/25/22 7:56 PM, Peter Nyikos wrote:
    On Wednesday, October 26, 2022 at 4:30:33 PM UTC-4, John Harshman wrote: >>>> On 10/26/22 12:57 PM, Peter Nyikos wrote:
    On Friday, October 21, 2022 at 10:14:35 PM UTC-4, John Harshman wrote: >>>>>> On 10/21/22 1:36 PM, Peter Nyikos wrote:
    On Friday, October 21, 2022 at 3:24:55 PM UTC-4, John Harshman wrote: >>>>>>>> On 10/26/22 12:57 PM, Peter Nyikos wrote:

    there are at least three bird lineages
    surviving the K/P extinction.

    Again, at least 7: paleognaths, neoaves, galliforms, anhimids, >>>>>>>> presbyornithids, anseranatidae, anatidae.

    Which bird did you have in mind from neoaves?

    It's worth noting that the last 5 groups you list are all in Galloanserae.

    And the last four are generally believed to be in Anseriformes ("waterfowl").

    "Generally believed" is a very weak statement, and I'm not sure why you
    put it that way. Yes, they are anseriforms.

    If, as is generally believed, the first split in Neornithes is between
    paleognaths and Galloanserae, you are assuming below that
    Neoaves and Galloanserae are sister groups, right?

    You don't answer this, but I soon figured out that I was right here.

    You are partly right and partly wrong. The first part of your sentence
    is wrong but the second is right, and in fact the two parts are
    contradictory.

    It's all implied by the tree if we suppose there are Cretaceous
    presbyornithids and that presbyornithids are the sister group of
    anatids. I wasn't referring to any other fossils.

    The problem with that, as we saw in the first reply, is
    that the claim of there being Cretaceous presbyornithids
    is shaky. You need several generally accepted Cretaceous
    members of Galloanserae to make the claim that there
    were 7 lineages.

    No, you only need one, as long as it's the right one. Either a
    presbyornithid or an anatid would do.

    You are assuming some other features of the phylogenetic tree
    of the order Anseriformes ("waterfowl") that you aren't
    identifying here. What are they?

    Just the usual (and well-confirmed) relationships. Are you in any doubt?

    Here is a claim of a Cretaceous presbyornithid:

    https://www.ncbi.nlm.nih.gov/pmc/articles/PMC4785986/

    It sounds rather tentative, so not too encouraging of "only need one":

    "Although screamers (Anhimidae) may have evolved from presbyornithid-like birds, the uncertainty as to whether skeletal features shared by presbyornithids and anhimids are plesiomorphic within Anseriformes or indicative of a close relationship between
    the two cannot yet be resolved. In any case, recognition of W. tedfordi as a presbyornithid calls for a reassessment of the phylogenetic position of Presbyornithidae within Anseriformes."

    Sounds like your hypothesis is much more easily falsifiable than the hypothesis that a
    species of *Merychippus* is directly ancestral to *Equus* in the horse family.

    ??

    One might also add Gastornithidae and Pelagornithidae, though I'm not
    sure whether those are both separate branches.

    Ah, one of my favorite bird families, those pelagornithids ["pseudotoothed birds"].
    The biggest one of all, with a wingspread over 20 feet, was found right here in South Carolina, while clearing land for the Charleston international airport.


    However, I saw no candidates for either family having Cretaceous members.
    And whether they are separate branches or the same seems irrelevant.
    You really have your work cut out for you to find an indirect phylogenetic tree way to make them survivors of the K-P extinction.

    You still don't seem to understand how this works. If we have a
    Cretaceous example of a particular branch, all nodes below that branch
    must also exist at that time. Given a Cretaceous presbyornithid, and
    give tnat Presbyornithidae is the sister group of Anatidae, there must
    be Cretaceous gastornithids and/or (depending on the tree)
    pelagornithids, and anseranatids, anhimids, neoavians, and paleognaths.

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From Peter Nyikos@21:1/5 to John Harshman on Wed Nov 30 14:10:43 2022
    On Tuesday, November 29, 2022 at 9:19:13 AM UTC-5, John Harshman wrote:
    On 11/28/22 7:13 PM, Peter Nyikos wrote:
    On Sunday, November 27, 2022 at 1:07:11 AM UTC-5, John Harshman wrote:
    On 11/25/22 7:56 PM, Peter Nyikos wrote:
    On Wednesday, October 26, 2022 at 4:30:33 PM UTC-4, John Harshman wrote: >>>> On 10/26/22 12:57 PM, Peter Nyikos wrote:
    On Friday, October 21, 2022 at 10:14:35 PM UTC-4, John Harshman wrote: >>>>>> On 10/21/22 1:36 PM, Peter Nyikos wrote:
    On Friday, October 21, 2022 at 3:24:55 PM UTC-4, John Harshman wrote:
    On 10/26/22 12:57 PM, Peter Nyikos wrote:

    there are at least three bird lineages
    surviving the K/P extinction.

    Again, at least 7: paleognaths, neoaves, galliforms, anhimids, >>>>>>>> presbyornithids, anseranatidae, anatidae.

    Which bird did you have in mind from neoaves?

    It's worth noting that the last 5 groups you list are all in Galloanserae.

    And the last four are generally believed to be in Anseriformes ("waterfowl").

    "Generally believed" is a very weak statement, and I'm not sure why you
    put it that way.

    Because Gerald Mayr, on p. 115 of his 2017 book, had some doubts about presbyornithids:

    "Unfortunately, an assessment of the affinities of presbyornithids is hampered by the poorly known morphology of the skull, which has not yet been studied in detail, despite the availability of numerous well-preserved specimens. The quadrate, the only
    skull bone of which more detailed descriptions exist, exhibits several presumably plesimorphic characteristics, which suggest a position of presbyornithids outside crown group Anseriformes [Elzanowski and Stidham 2010]."


    Yes, they are anseriforms.

    That's a very strong statement. :)


    If, as is generally believed, the first split in Neornithes is between >>> paleognaths and Galloanserae, you are assuming below that
    Neoaves and Galloanserae are sister groups, right?

    You don't answer this, but I soon figured out that I was right here.

    You are partly right and partly wrong. The first part of your sentence
    is wrong

    Sorry, I was absent-minded. I meant "neognaths" instead
    of Galloanserae in the first part.

    However, the possibility that there is a poorly resolved trichotomy
    between the three groups occurred to me just now. Recall the example of {theropods, sauropods, ornithischians} at the base of Dinosauria.
    Is it different with the base of Neornithes?


    It's all implied by the tree if we suppose there are Cretaceous >>>>>> presbyornithids and that presbyornithids are the sister group of >>>>>> anatids. I wasn't referring to any other fossils.

    The problem with that, as we saw in the first reply, is
    that the claim of there being Cretaceous presbyornithids
    is shaky. You need several generally accepted Cretaceous
    members of Galloanserae to make the claim that there
    were 7 lineages.

    No, you only need one, as long as it's the right one. Either a
    presbyornithid or an anatid would do.

    You are assuming some other features of the phylogenetic tree
    of the order Anseriformes ("waterfowl") that you aren't
    identifying here. What are they?

    Just the usual (and well-confirmed) relationships.

    Why so tight-lipped? I suppose you are referring to the tree of Anseriformes
    in Wikipedia, but since when is Wikipedia an infallible guide to phylogeny?

    Are you in any doubt?

    Yes, whenever you are as tight-lipped as this. Besides, it's bad form:
    what is sci.bio.paleontology for, if not for liberal conveying
    of information among participants? I say "among" instead of "between"
    because I think other regulars can benefit from a more informative
    answer to the question I asked. That's why I wrote "waterfowl" after "Anseriformes" because there might be some readers who are unfamiliar
    with the technical designations of bird orders.


    CONCLUDED in next reply, to be done not long after I see
    that this one has posted.


    Peter Nyikos
    Professor, Dept. of Mathematics -- standard disclaimer--
    Univ. of So. Carolina in Columbia
    http://people.math.sc.edu/nyikos

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From Peter Nyikos@21:1/5 to John Harshman on Wed Nov 30 15:23:20 2022
    On Tuesday, November 29, 2022 at 9:19:13 AM UTC-5, John Harshman wrote:
    On 11/28/22 7:13 PM, Peter Nyikos wrote:
    On Sunday, November 27, 2022 at 1:07:11 AM UTC-5, John Harshman wrote:
    On 11/25/22 7:56 PM, Peter Nyikos wrote:
    On Wednesday, October 26, 2022 at 4:30:33 PM UTC-4, John Harshman wrote: >>>> On 10/26/22 12:57 PM, Peter Nyikos wrote:
    On Friday, October 21, 2022 at 10:14:35 PM UTC-4, John Harshman wrote: >>>>>> On 10/21/22 1:36 PM, Peter Nyikos wrote:
    On Friday, October 21, 2022 at 3:24:55 PM UTC-4, John Harshman wrote:
    On 10/26/22 12:57 PM, Peter Nyikos wrote:

    there are at least three bird lineages
    surviving the K/P extinction.

    Again, at least 7: paleognaths, neoaves, galliforms, anhimids, >>>>>>>> presbyornithids, anseranatidae, anatidae.


    <snip to get to where I left off in my first reply>


    Here is a claim of a Cretaceous presbyornithid:

    https://www.ncbi.nlm.nih.gov/pmc/articles/PMC4785986/

    It sounds rather tentative, so not too encouraging of "only need one":

    "Although screamers (Anhimidae) may have evolved from presbyornithid-like birds, the uncertainty as to whether skeletal features shared by presbyornithids and anhimids are plesiomorphic within Anseriformes or indicative of a close relationship
    between the two cannot yet be resolved. In any case, recognition of W. tedfordi as a presbyornithid calls for a reassessment of the phylogenetic position of Presbyornithidae within Anseriformes."

    Sounds like your hypothesis is much more easily falsifiable than the hypothesis that a
    species of *Merychippus* is directly ancestral to *Equus* in the horse family.

    ??

    Come on, John, the general theme of direct ancestry
    has been a perennial bone of contention between us.
    If you disagree in this instance, please explain your disagreement.


    One might also add Gastornithidae and Pelagornithidae, though I'm not
    sure whether those are both separate branches.

    Ah, one of my favorite bird families, those pelagornithids ["pseudotoothed birds"].
    The biggest one of all, with a wingspread over 20 feet, was found right here
    in South Carolina, while clearing land for the Charleston international airport.

    I'm glad you mentioned them. I found out a fascinating piece of information from the
    Wikipedia entry on them just now.

    "They were the dominant seabirds of most oceans throughout most of the Cenozoic, and modern humans apparently missed encountering them only by a tiny measure of evolutionary time: the last known pelagornithids were contemporaries of Homo habilis and the
    beginning of the history of technology.[3]"

    So near and yet so far. :(


    However, I saw no candidates for either family having Cretaceous members. And whether they are separate branches or the same seems irrelevant.
    You really have your work cut out for you to find an indirect phylogenetic tree way to make them survivors of the K-P extinction.

    You still don't seem to understand how this works.

    I don't understand why you say this. You are ignoring the fact that Gastornis is considered by some to be within crown Anseriformes.

    As for pelagornithids, the Wikipedia entry tentatively put them into Galloanserae,
    but then treated us to a long discussion that reads like a technical review paper
    of various proposed possibilities, including some "outliers" in Neoaves.
    .

    If we have a
    Cretaceous example of a particular branch, all nodes below that branch
    must also exist at that time. Given a Cretaceous presbyornithid, and
    give tnat Presbyornithidae is the sister group of Anatidae, there must
    be Cretaceous gastornithids and/or (depending on the tree)
    pelagornithids, and anseranatids, anhimids, neoavians, and paleognaths.

    Too many assumptions here, in the light of my latest comments.
    Add to them the possibility that Mayr gave [see my first reply to your post], that presbyornithids are outside Anseriformes, and you could be down to paleognaths, neoaves, galliforms, presbyornithids, and a cladogenesis
    at some Paleocene member of total Anseriformes after anagenesis
    (or evolutionary stasis) from where presbyornithids split off from from Anseriformes.

    Five lineages, not seven.


    Peter Nyikos
    Professor, Dept. of Mathematics
    Univ. of So. Carolina -- standard disclaimer-- http://people.math.sc.edu/nyikos

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From John Harshman@21:1/5 to Peter Nyikos on Wed Nov 30 16:49:31 2022
    On 11/30/22 3:23 PM, Peter Nyikos wrote:
    On Tuesday, November 29, 2022 at 9:19:13 AM UTC-5, John Harshman wrote:
    On 11/28/22 7:13 PM, Peter Nyikos wrote:
    On Sunday, November 27, 2022 at 1:07:11 AM UTC-5, John Harshman wrote:
    On 11/25/22 7:56 PM, Peter Nyikos wrote:
    On Wednesday, October 26, 2022 at 4:30:33 PM UTC-4, John Harshman wrote: >>>>>> On 10/26/22 12:57 PM, Peter Nyikos wrote:
    On Friday, October 21, 2022 at 10:14:35 PM UTC-4, John Harshman wrote: >>>>>>>> On 10/21/22 1:36 PM, Peter Nyikos wrote:
    On Friday, October 21, 2022 at 3:24:55 PM UTC-4, John Harshman wrote: >>>>>>>>>> On 10/26/22 12:57 PM, Peter Nyikos wrote:

    there are at least three bird lineages
    surviving the K/P extinction.

    Again, at least 7: paleognaths, neoaves, galliforms, anhimids, >>>>>>>>>> presbyornithids, anseranatidae, anatidae.


    <snip to get to where I left off in my first reply>


    Here is a claim of a Cretaceous presbyornithid:

    https://www.ncbi.nlm.nih.gov/pmc/articles/PMC4785986/

    It sounds rather tentative, so not too encouraging of "only need one":

    "Although screamers (Anhimidae) may have evolved from presbyornithid-like birds, the uncertainty as to whether skeletal features shared by presbyornithids and anhimids are plesiomorphic within Anseriformes or indicative of a close relationship
    between the two cannot yet be resolved. In any case, recognition of W. tedfordi as a presbyornithid calls for a reassessment of the phylogenetic position of Presbyornithidae within Anseriformes."

    Sounds like your hypothesis is much more easily falsifiable than the hypothesis that a
    species of *Merychippus* is directly ancestral to *Equus* in the horse family.

    ??

    Come on, John, the general theme of direct ancestry
    has been a perennial bone of contention between us.
    If you disagree in this instance, please explain your disagreement.

    I truly don't know what you were trying to hint at there, so I don't
    know whether I agree.

    One might also add Gastornithidae and Pelagornithidae, though I'm not
    sure whether those are both separate branches.

    Ah, one of my favorite bird families, those pelagornithids ["pseudotoothed birds"].
    The biggest one of all, with a wingspread over 20 feet, was found right here
    in South Carolina, while clearing land for the Charleston international airport.

    I'm glad you mentioned them. I found out a fascinating piece of information from the
    Wikipedia entry on them just now.

    "They were the dominant seabirds of most oceans throughout most of the Cenozoic, and modern humans apparently missed encountering them only by a tiny measure of evolutionary time: the last known pelagornithids were contemporaries of Homo habilis and
    the beginning of the history of technology.[3]"

    So near and yet so far. :(

    Similar story for phorusrhacids.

    However, I saw no candidates for either family having Cretaceous members. >>> And whether they are separate branches or the same seems irrelevant.
    You really have your work cut out for you to find an indirect phylogenetic >>> tree way to make them survivors of the K-P extinction.

    You still don't seem to understand how this works.

    I don't understand why you say this. You are ignoring the fact that Gastornis is considered by some to be within crown Anseriformes.

    I'm not aware of that. Who?

    As for pelagornithids, the Wikipedia entry tentatively put them into Galloanserae,
    but then treated us to a long discussion that reads like a technical review paper
    of various proposed possibilities, including some "outliers" in Neoaves.

    Sounds like it was written by a very old specialist. Better to look for
    pubs with actual phylogenetic analyses in them, none of which I have
    off-hand.

    >If we have a
    Cretaceous example of a particular branch, all nodes below that branch
    must also exist at that time. Given a Cretaceous presbyornithid, and
    give tnat Presbyornithidae is the sister group of Anatidae, there must
    be Cretaceous gastornithids and/or (depending on the tree)
    pelagornithids, and anseranatids, anhimids, neoavians, and paleognaths.

    Too many assumptions here, in the light of my latest comments.
    Add to them the possibility that Mayr gave [see my first reply to your post], that presbyornithids are outside Anseriformes, and you could be down to paleognaths, neoaves, galliforms, presbyornithids, and a cladogenesis
    at some Paleocene member of total Anseriformes after anagenesis
    (or evolutionary stasis) from where presbyornithids split off from from Anseriformes.

    Five lineages, not seven.

    Yes, that's possible, though whatever you are trying to say regarding anagenesis is unclear. Just "anseriforms" would have been simpler and
    more correct. So we're agreed that you don't need more than one
    Cretaceous fossil to infer the Cretaceous existence of several other
    groups. That's the point I was trying to get across. The meaning of
    "several" depends on just where presbyornithids end up.

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From John Harshman@21:1/5 to Peter Nyikos on Wed Nov 30 16:39:26 2022
    On 11/30/22 2:10 PM, Peter Nyikos wrote:
    On Tuesday, November 29, 2022 at 9:19:13 AM UTC-5, John Harshman wrote:
    On 11/28/22 7:13 PM, Peter Nyikos wrote:
    On Sunday, November 27, 2022 at 1:07:11 AM UTC-5, John Harshman wrote:
    On 11/25/22 7:56 PM, Peter Nyikos wrote:
    On Wednesday, October 26, 2022 at 4:30:33 PM UTC-4, John Harshman wrote: >>>>>> On 10/26/22 12:57 PM, Peter Nyikos wrote:
    On Friday, October 21, 2022 at 10:14:35 PM UTC-4, John Harshman wrote: >>>>>>>> On 10/21/22 1:36 PM, Peter Nyikos wrote:
    On Friday, October 21, 2022 at 3:24:55 PM UTC-4, John Harshman wrote: >>>>>>>>>> On 10/26/22 12:57 PM, Peter Nyikos wrote:

    there are at least three bird lineages
    surviving the K/P extinction.

    Again, at least 7: paleognaths, neoaves, galliforms, anhimids, >>>>>>>>>> presbyornithids, anseranatidae, anatidae.

    Which bird did you have in mind from neoaves?

    It's worth noting that the last 5 groups you list are all in Galloanserae.

    And the last four are generally believed to be in Anseriformes ("waterfowl").

    "Generally believed" is a very weak statement, and I'm not sure why you
    put it that way.

    Because Gerald Mayr, on p. 115 of his 2017 book, had some doubts about presbyornithids:

    "Unfortunately, an assessment of the affinities of presbyornithids is hampered by the poorly known morphology of the skull, which has not yet been studied in detail, despite the availability of numerous well-preserved specimens. The quadrate, the only
    skull bone of which more detailed descriptions exist, exhibits several presumably plesimorphic characteristics, which suggest a position of presbyornithids outside crown group Anseriformes [Elzanowski and Stidham 2010]."


    Yes, they are anseriforms.

    That's a very strong statement. :)

    Well, they have duck bills. The skull may not have been described
    completely but it's been well illustrated by Olson & Feduccia. I'll have
    to see what Elzanowski & Stidham have to say.

    If, as is generally believed, the first split in Neornithes is between >>>>> paleognaths and Galloanserae, you are assuming below that
    Neoaves and Galloanserae are sister groups, right?

    You don't answer this, but I soon figured out that I was right here.

    You are partly right and partly wrong. The first part of your sentence
    is wrong

    Sorry, I was absent-minded. I meant "neognaths" instead
    of Galloanserae in the first part.

    However, the possibility that there is a poorly resolved trichotomy
    between the three groups occurred to me just now. Recall the example of {theropods, sauropods, ornithischians} at the base of Dinosauria.
    Is it different with the base of Neornithes?

    Yes, very different. There is zero doubt on that particular pair of nodes.

    It's all implied by the tree if we suppose there are Cretaceous >>>>>>>> presbyornithids and that presbyornithids are the sister group of >>>>>>>> anatids. I wasn't referring to any other fossils.

    The problem with that, as we saw in the first reply, is
    that the claim of there being Cretaceous presbyornithids
    is shaky. You need several generally accepted Cretaceous
    members of Galloanserae to make the claim that there
    were 7 lineages.

    No, you only need one, as long as it's the right one. Either a
    presbyornithid or an anatid would do.

    You are assuming some other features of the phylogenetic tree
    of the order Anseriformes ("waterfowl") that you aren't
    identifying here. What are they?

    Just the usual (and well-confirmed) relationships.

    Why so tight-lipped? I suppose you are referring to the tree of Anseriformes in Wikipedia, but since when is Wikipedia an infallible guide to phylogeny?

    I am not referring to the Wikipedia tree. The basal nodes of the
    anseriform tree (excluding Presbyornis) have been confirmed by numerous molecular and morphological analyses. It's (Animidae,(Anseranatidae, Anatidae)). Again, there is no doubt on that point.

    For example, see Hackett S.J., Kimball R.T., Reddy S., Bowie R.C.K.,
    Braun E.L., Braun M.J., Chojnowski J.L., Cox W.A., Han K.-L., Harshman
    J., Huddleston C.J., Marks B.D., Miglia K.J., Moore W.A., Sheldon F.H., Steadman D.W., Witt C.C., Yuri T. A phylogenomic study of birds reveals
    their evolutionary history. Science 2008; 320:1763-1768.

    Looking at the Wikipedia tree, I see that it's correct on everything up
    until Plectropterus.

    The basal nodes of Galliformes are equally well resolved.

    Are you in any doubt?

    Yes, whenever you are as tight-lipped as this. Besides, it's bad form:
    what is sci.bio.paleontology for, if not for liberal conveying
    of information among participants? I say "among" instead of "between"
    because I think other regulars can benefit from a more informative
    answer to the question I asked. That's why I wrote "waterfowl" after "Anseriformes" because there might be some readers who are unfamiliar
    with the technical designations of bird orders.


    CONCLUDED in next reply, to be done not long after I see
    that this one has posted.


    Peter Nyikos
    Professor, Dept. of Mathematics -- standard disclaimer--
    Univ. of So. Carolina in Columbia
    http://people.math.sc.edu/nyikos

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From Daud Deden@21:1/5 to peter2...@gmail.com on Thu Dec 1 22:18:03 2022
    On Wednesday, November 9, 2022 at 9:14:07 PM UTC-5, peter2...@gmail.com wrote:
    The Great K-T extinction spelled the doom of the giant sea reptiles, all dinosaurs
    (unless you count birds as dinosaurs), and many other groups of vertebrates and invertebrates.
    In fact, the damage caused by the asteroid that hit Yucatan was so great that no animal over 50 kilos is known to have survived.

    But some groups of birds and mammals did survive, including the remote ancestors
    of the four groups that have survived to this day: the neornithine birds and three groups of mammals:
    the placentals, the marsupials, and the monotremes.

    To keep this OP reasonably short, I will deal with the two most familiar of the four, the placentals and the modern birds.

    A 2013 _Science_ article [1] led to the general acceptance of the hypothesis that all living placentals
    have descended from a common ancestor that lived after the great K-T disaster.

    However, there are closely related eutherians [2] that also survived for a while, among
    whose descendants were the pantodonts, including *Coryphodon*, which was a bit larger than a tapir; and the tillodonts and taeniodonts. These groups died out during the Eocene, which ended almost halfway between the K-T disaster and our time.

    The case of birds is generally believed to be quite different: EVERY bird, living
    or extinct, that survived the K-T disaster is believed to be descended from the
    last common ancestor of living birds. However, there are some articles which hypothesize that there is one exception, a group of birds called the lithornids.
    More about them in my next post to this thread, some time tomorrow.

    [1] Maureen A. O'Leary et. al., "The Placental Mammal Ancestor and the Post–K-Pg Radiation of Placentals". Science. 339 (6120): 662–667.

    [2] Eutherians are the mammals, extant and extinct, that are more closely related to placentals than to marsupials.
    Similarly, metatherians are the mammals that are more closely related to marsupials than to placentals.


    Peter Nyikos
    Professor, Dept. of Mathematics -- standard disclaimer--
    University of South Carolina
    http://people.math.sc.edu/nyikos

    PS The "Why?" at the end of the thread title is two-edged: why did some survive,
    and why did others, including some similar-seeming ones, perish?
    -
    Not sure where to put this: https://www.nature.com/articles/d41586-022-04181-7

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From erik simpson@21:1/5 to daud....@gmail.com on Fri Dec 2 08:52:33 2022
    On Thursday, December 1, 2022 at 10:18:04 PM UTC-8, daud....@gmail.com wrote:
    On Wednesday, November 9, 2022 at 9:14:07 PM UTC-5, peter2...@gmail.com wrote:
    The Great K-T extinction spelled the doom of the giant sea reptiles, all dinosaurs
    (unless you count birds as dinosaurs), and many other groups of vertebrates and invertebrates.
    In fact, the damage caused by the asteroid that hit Yucatan was so great that no animal over 50 kilos is known to have survived.

    But some groups of birds and mammals did survive, including the remote ancestors
    of the four groups that have survived to this day: the neornithine birds and three groups of mammals:
    the placentals, the marsupials, and the monotremes.

    To keep this OP reasonably short, I will deal with the two most familiar of the four, the placentals and the modern birds.

    A 2013 _Science_ article [1] led to the general acceptance of the hypothesis that all living placentals
    have descended from a common ancestor that lived after the great K-T disaster.

    However, there are closely related eutherians [2] that also survived for a while, among
    whose descendants were the pantodonts, including *Coryphodon*, which was a bit larger than a tapir; and the tillodonts and taeniodonts. These groups died out during the Eocene, which ended almost halfway between the K-T disaster and our time.

    The case of birds is generally believed to be quite different: EVERY bird, living
    or extinct, that survived the K-T disaster is believed to be descended from the
    last common ancestor of living birds. However, there are some articles which
    hypothesize that there is one exception, a group of birds called the lithornids.
    More about them in my next post to this thread, some time tomorrow.

    [1] Maureen A. O'Leary et. al., "The Placental Mammal Ancestor and the Post–K-Pg Radiation of Placentals". Science. 339 (6120): 662–667.

    [2] Eutherians are the mammals, extant and extinct, that are more closely related to placentals than to marsupials.
    Similarly, metatherians are the mammals that are more closely related to marsupials than to placentals.


    Peter Nyikos
    Professor, Dept. of Mathematics -- standard disclaimer--
    University of South Carolina
    http://people.math.sc.edu/nyikos

    PS The "Why?" at the end of the thread title is two-edged: why did some survive,
    and why did others, including some similar-seeming ones, perish?
    -
    Not sure where to put this: https://www.nature.com/articles/d41586-022-04181-7
    Good catch! The actual article: https://www.nature.com/articles/s41586-022-05445-y

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From Peter Nyikos@21:1/5 to John Harshman on Fri Dec 2 11:25:44 2022
    On Wednesday, November 30, 2022 at 7:49:38 PM UTC-5, John Harshman wrote:
    On 11/30/22 3:23 PM, Peter Nyikos wrote:
    On Tuesday, November 29, 2022 at 9:19:13 AM UTC-5, John Harshman wrote:
    On 11/28/22 7:13 PM, Peter Nyikos wrote:
    On Sunday, November 27, 2022 at 1:07:11 AM UTC-5, John Harshman wrote: >>>> On 11/25/22 7:56 PM, Peter Nyikos wrote:
    On Wednesday, October 26, 2022 at 4:30:33 PM UTC-4, John Harshman wrote:
    On 10/26/22 12:57 PM, Peter Nyikos wrote:
    On Friday, October 21, 2022 at 10:14:35 PM UTC-4, John Harshman wrote:
    On 10/21/22 1:36 PM, Peter Nyikos wrote:
    On Friday, October 21, 2022 at 3:24:55 PM UTC-4, John Harshman wrote:
    On 10/26/22 12:57 PM, Peter Nyikos wrote:

    there are at least three bird lineages
    surviving the K/P extinction.

    Again, at least 7: paleognaths, neoaves, galliforms, anhimids, >>>>>>>>>> presbyornithids, anseranatidae, anatidae.


    <snip to get to where I left off in my first reply>


    Here is a claim of a Cretaceous presbyornithid:

    https://www.ncbi.nlm.nih.gov/pmc/articles/PMC4785986/

    It sounds rather tentative, so not too encouraging of "only need one": >>>
    "Although screamers (Anhimidae) may have evolved from presbyornithid-like birds, the uncertainty as to whether skeletal features shared by presbyornithids and anhimids are plesiomorphic within Anseriformes or indicative of a close relationship
    between the two cannot yet be resolved. In any case, recognition of W. tedfordi as a presbyornithid calls for a reassessment of the phylogenetic position of Presbyornithidae within Anseriformes."

    Sounds like your hypothesis is much more easily falsifiable than the hypothesis that a
    species of *Merychippus* is directly ancestral to *Equus* in the horse family.

    ??

    Come on, John, the general theme of direct ancestry
    has been a perennial bone of contention between us.
    If you disagree in this instance, please explain your disagreement.

    I truly don't know what you were trying to hint at there, so I don't
    know whether I agree.

    I was not "hinting". What part of "your hypothesis is much more easily falsifiable than"
    don't you understand? We have essentially complete fossils of Merychippus and can study Equus to our heart's content.

    Compare that to the evidence that *Teviornis gobiensis* was a Cretaceous presbyornithid.
    Mayr's 2017 book doesn't even mention the genus nor the 2016 paper that we have been
    discussing above. Neither does Wikipedia, which only cites two papers on the genus,
    in 2002 and 2004. Do you know of any citations of it in the peer-reviewed literature?
    Or any more recent evidence for its tentative conclusion?

    It's an awfully thin peg to hang 7 separate lineages on.

    One might also add Gastornithidae and Pelagornithidae, though I'm not >>>> sure whether those are both separate branches.

    Ah, one of my favorite bird families, those pelagornithids ["pseudotoothed birds"].
    The biggest one of all, with a wingspread over 20 feet, was found right here
    in South Carolina, while clearing land for the Charleston international airport.

    I'm glad you mentioned them. I found out a fascinating piece of information from the
    Wikipedia entry on them just now.

    "They were the dominant seabirds of most oceans throughout most of the Cenozoic, and modern humans apparently missed encountering them only by a tiny measure of evolutionary time: the last known pelagornithids were contemporaries of Homo habilis and
    the beginning of the history of technology.[3]"

    So near and yet so far. :(

    Similar story for phorusrhacids.

    Even more so: put Homo erectus there. However, the much more recent claims of ca.100 kya
    and even 18 kya are not well supported according to the entry on them.
    The summary in the box at upper right should have a question mark added after 0.1 Ma.
    Another case of discordant statements on the same Wiki page.


    However, I saw no candidates for either family having Cretaceous members.
    And whether they are separate branches or the same seems irrelevant.
    You really have your work cut out for you to find an indirect phylogenetic
    tree way to make them survivors of the K-P extinction.

    You still don't seem to understand how this works.

    I don't understand why you say this. You are ignoring the fact that Gastornis
    is considered by some to be within crown Anseriformes.

    I'm not aware of that. Who?

    Read the Wikipedia entry.

    As for pelagornithids, the Wikipedia entry tentatively put them into Galloanserae,
    but then treated us to a long discussion that reads like a technical review paper
    of various proposed possibilities, including some "outliers" in Neoaves.

    Sounds like it was written by a very old specialist. Better to look for
    pubs with actual phylogenetic analyses in them, none of which I have off-hand.

    Please let us know when you find any. But keep in mind that the
    reviewer cites a number of them already.


    If we have a
    Cretaceous example of a particular branch, all nodes below that branch
    must also exist at that time. Given a Cretaceous presbyornithid, and
    give tnat Presbyornithidae is the sister group of Anatidae, there must
    be Cretaceous gastornithids and/or (depending on the tree)
    pelagornithids, and anseranatids, anhimids, neoavians, and paleognaths.

    Too many assumptions here, in the light of my latest comments.
    Add to them the possibility that Mayr gave [see my first reply to your post],
    that presbyornithids are outside Anseriformes, and you could be down to paleognaths, neoaves, galliforms, presbyornithids, and a cladogenesis
    at some Paleocene member of total Anseriformes after anagenesis
    (or evolutionary stasis) from where presbyornithids split off from from Anseriformes.

    Five lineages, not seven.

    Yes, that's possible, though whatever you are trying to say regarding anagenesis is unclear.

    One or more speciations without branching. The old species may
    linger on for a while, but in the absence of fossils for side branches,
    there is no evidence for cladogenesis.

    We're talking about lineages, not individual taxa.


    Just "anseriforms" would have been simpler and
    more correct.

    How so? doesn't it mean the same thing as Anseriformes?


    So we're agreed that you don't need more than one
    Cretaceous fossil to infer the Cretaceous existence of several other
    groups.

    Of course, but the fossil involved in this case is fragmentary.


    I've asked Trevor Worthy last night (my time; this morning his time)
    in email about how the 2016 paper has fared since then, and I hope he answers before long.


    Peter Nyikos
    Professor, Dept. of Mathematics -- standard disclaimer--
    U. of South Carolina at Columbia
    http://people.math.sc.edu/nyikos

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From Peter Nyikos@21:1/5 to daud....@gmail.com on Fri Dec 2 16:11:45 2022
    On Friday, December 2, 2022 at 1:18:04 AM UTC-5, daud....@gmail.com wrote:
    On Wednesday, November 9, 2022 at 9:14:07 PM UTC-5, peter2...@gmail.com wrote:

    PS The "Why?" at the end of the thread title is two-edged: why did some survive,
    and why did others, including some similar-seeming ones, perish?
    -
    Not sure where to put this:

    "67-million-year-old fossil upends bird evolutionary tree"
    https://www.nature.com/articles/d41586-022-04181-7

    It's perfect for this thread. It speaks to both halves of the title,
    by the way it upends the evolutionary tree and has a datum for "Why":
    a bird that did not survive and may have perished
    in the asteroid-related disaster. I'll be speaking about
    this "Why" today and save the details about the upending on
    Monday. It is relevant to which lineages of birds survived the disaster.


    Before I go on: even if you are rushed for time, if the title of the article gives a good hint for what it's about, it's worth a few seconds
    to highlight, copy, and paste. Here, with such a dramatic title,
    it's very worthwhile. So I put it in for you.

    Frosting on the cake: you've got a real scoop as far as s.b.p. is concerned.
    As they used to say before the internet: it's hot off the presses.
    It's dated 30 November 2022, day before yesterday.

    Even more: although the title is dramatic, there's no hype.
    I'm fortunate to have learned enough details about the base
    of the neornithine tree in just the last month to fully appreciate this.
    I'll have more to say on this on Monday, after my usual weekend posting break.


    For now, back to that datum. Here is the research article
    for which the one you've scooped is the announcement for the general public.

    Cretaceous ornithurine supports a neognathous crown bird ancestor https://www.nature.com/articles/s41586-022-05445-y

    Here is where it speaks to whether its extinction was due to the K-T disaster, and why:

    " thus far, *Janavis* constitutes the only confidently identified and non-fragmentary Maastrichtian ichthyornithine known, and its large size and presumed extinction at the end-Cretaceous are congruent with hypothesized selection favouring relatively
    small-bodied birds across the Cretaceous–Paleogene (K–Pg) boundary28,29."

    Not just Maastrichtian, but Latest Maastrichtian. [1] As for the talk about size,
    I keep reading that no animal over 50 kilos is known to have survived. I also believe
    that most animals over 10 kilos had serious trouble foraging for enough food to keep alive
    after the asteroid struck. *Janavis* is nowhere near this big -- it's estimated to
    have been between 1 and 1.5 kilos -- but it's still a lot bigger than its iconic relative, *Ichthyornis*.
    Moreover:
    "Indeed, Janavis greatly outweighs the only other co-occurring avialan known from the Maastricht Formation: the early neornithine Asteriornis maastrichtensis, the holotype of which is estimated to have weighed in the range of 395 g (ref. 27)."


    [1] The Maastrichtian is the last big division of the Cretaceous. For a decade or two, I'd get goose
    bumps every time I came across the word "Maastrichthian" because I knew what it portended.
    If the link Erik gave us works for you, take a look at Fig.1 [it can be magnified] and look to the
    right to see *Janavis* with a line to it that ends at the very end of the Maastrichthian.

    Appropriately, the holotype is in Natuurhistorisch Museum Maastricht.

    I close this post with a comment about the title of the research paper, "Cretaceous ornithurine supports a neognathous crown bird ancestor"

    Although it may seem prosaic compared to the one for the paper you scooped,
    it actually arouses my interest more because of all the information it conveys.

    Orniturine birds include neornithine (crown) birds, ichthyornithids, hesperornithids,
    and a handful of birds that are, with one exception, less closely related to crown birds.
    So *Janavis* can be expected to provide valuable information about the base
    of the crown birds. The rest of the title speaks volumes about the "upending" of the base, and with it the tree, to a trained eye, as I'll explain on Monday unless someone beats me to it.


    Peter Nyikos
    Professor, Dept. of Mathematics -- standard disclaimer--
    U. of South Carolina in Columbia
    http://people.math.sc.edu/nyikos

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From John Harshman@21:1/5 to Daud Deden on Fri Dec 2 16:52:35 2022
    On 12/1/22 10:18 PM, Daud Deden wrote:
    On Wednesday, November 9, 2022 at 9:14:07 PM UTC-5, peter2...@gmail.com wrote:
    The Great K-T extinction spelled the doom of the giant sea reptiles, all dinosaurs
    (unless you count birds as dinosaurs), and many other groups of vertebrates and invertebrates.
    In fact, the damage caused by the asteroid that hit Yucatan was so great
    that no animal over 50 kilos is known to have survived.

    But some groups of birds and mammals did survive, including the remote
    ancestors
    of the four groups that have survived to this day: the neornithine birds and three groups of mammals:
    the placentals, the marsupials, and the monotremes.

    To keep this OP reasonably short, I will deal with the two most familiar of the four, the placentals and the modern birds.

    A 2013 _Science_ article [1] led to the general acceptance of the hypothesis that all living placentals
    have descended from a common ancestor that lived after the great K-T disaster.

    However, there are closely related eutherians [2] that also survived for a while, among
    whose descendants were the pantodonts, including *Coryphodon*, which was a bit larger than a tapir; and the tillodonts and taeniodonts. These groups died out during the Eocene, which ended almost halfway between the K-T disaster and our time.

    The case of birds is generally believed to be quite different: EVERY bird, living
    or extinct, that survived the K-T disaster is believed to be descended from the
    last common ancestor of living birds. However, there are some articles which >> hypothesize that there is one exception, a group of birds called the lithornids.
    More about them in my next post to this thread, some time tomorrow.

    [1] Maureen A. O'Leary et. al., "The Placental Mammal Ancestor and the Post–K-Pg Radiation of Placentals". Science. 339 (6120): 662–667.

    [2] Eutherians are the mammals, extant and extinct, that are more closely related to placentals than to marsupials.
    Similarly, metatherians are the mammals that are more closely related to marsupials than to placentals.


    Peter Nyikos
    Professor, Dept. of Mathematics -- standard disclaimer--
    University of South Carolina
    http://people.math.sc.edu/nyikos

    PS The "Why?" at the end of the thread title is two-edged: why did some survive,
    and why did others, including some similar-seeming ones, perish?
    -
    Not sure where to put this: https://www.nature.com/articles/d41586-022-04181-7

    As usual in science journalism, the title (at least) is hype. This
    discovery has no effect on the evolutionary tree, just on the character
    state at the base of the tree. Paleognaths and neognaths are still each monophyletic. No change there.

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From Daud Deden@21:1/5 to John Harshman on Fri Dec 2 17:08:40 2022
    On Friday, December 2, 2022 at 7:52:41 PM UTC-5, John Harshman wrote:
    On 12/1/22 10:18 PM, Daud Deden wrote:
    On Wednesday, November 9, 2022 at 9:14:07 PM UTC-5, peter2...@gmail.com wrote:
    The Great K-T extinction spelled the doom of the giant sea reptiles, all dinosaurs
    (unless you count birds as dinosaurs), and many other groups of vertebrates and invertebrates.
    In fact, the damage caused by the asteroid that hit Yucatan was so great >> that no animal over 50 kilos is known to have survived.

    But some groups of birds and mammals did survive, including the remote
    ancestors
    of the four groups that have survived to this day: the neornithine birds and three groups of mammals:
    the placentals, the marsupials, and the monotremes.

    To keep this OP reasonably short, I will deal with the two most familiar of the four, the placentals and the modern birds.

    A 2013 _Science_ article [1] led to the general acceptance of the hypothesis that all living placentals
    have descended from a common ancestor that lived after the great K-T disaster.

    However, there are closely related eutherians [2] that also survived for a while, among
    whose descendants were the pantodonts, including *Coryphodon*, which was a bit larger than a tapir; and the tillodonts and taeniodonts. These groups died out during the Eocene, which ended almost halfway between the K-T disaster and our time.

    The case of birds is generally believed to be quite different: EVERY bird, living
    or extinct, that survived the K-T disaster is believed to be descended from the
    last common ancestor of living birds. However, there are some articles which
    hypothesize that there is one exception, a group of birds called the lithornids.
    More about them in my next post to this thread, some time tomorrow.

    [1] Maureen A. O'Leary et. al., "The Placental Mammal Ancestor and the Post–K-Pg Radiation of Placentals". Science. 339 (6120): 662–667.

    [2] Eutherians are the mammals, extant and extinct, that are more closely related to placentals than to marsupials.
    Similarly, metatherians are the mammals that are more closely related to marsupials than to placentals.


    Peter Nyikos
    Professor, Dept. of Mathematics -- standard disclaimer--
    University of South Carolina
    http://people.math.sc.edu/nyikos

    PS The "Why?" at the end of the thread title is two-edged: why did some survive,
    and why did others, including some similar-seeming ones, perish?
    -
    Not sure where to put this: https://www.nature.com/articles/d41586-022-04181-7
    As usual in science journalism, the title (at least) is hype. This
    discovery has no effect on the evolutionary tree, just on the character state at the base of the tree. Paleognaths and neognaths are still each monophyletic. No change there.

    OT streamlined theropod natovenator no wings https://www.nature.com/articles/s42003-022-04119-9

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From John Harshman@21:1/5 to Peter Nyikos on Fri Dec 2 16:43:48 2022
    On 12/2/22 11:25 AM, Peter Nyikos wrote:
    On Wednesday, November 30, 2022 at 7:49:38 PM UTC-5, John Harshman wrote:
    On 11/30/22 3:23 PM, Peter Nyikos wrote:
    On Tuesday, November 29, 2022 at 9:19:13 AM UTC-5, John Harshman wrote: >>>> On 11/28/22 7:13 PM, Peter Nyikos wrote:
    On Sunday, November 27, 2022 at 1:07:11 AM UTC-5, John Harshman wrote: >>>>>> On 11/25/22 7:56 PM, Peter Nyikos wrote:
    On Wednesday, October 26, 2022 at 4:30:33 PM UTC-4, John Harshman wrote:
    On 10/26/22 12:57 PM, Peter Nyikos wrote:
    On Friday, October 21, 2022 at 10:14:35 PM UTC-4, John Harshman wrote:
    On 10/21/22 1:36 PM, Peter Nyikos wrote:
    On Friday, October 21, 2022 at 3:24:55 PM UTC-4, John Harshman wrote:
    On 10/26/22 12:57 PM, Peter Nyikos wrote:

    there are at least three bird lineages
    surviving the K/P extinction.

    Again, at least 7: paleognaths, neoaves, galliforms, anhimids, >>>>>>>>>>>> presbyornithids, anseranatidae, anatidae.


    <snip to get to where I left off in my first reply>


    Here is a claim of a Cretaceous presbyornithid:

    https://www.ncbi.nlm.nih.gov/pmc/articles/PMC4785986/

    It sounds rather tentative, so not too encouraging of "only need one": >>>>>
    "Although screamers (Anhimidae) may have evolved from presbyornithid-like birds, the uncertainty as to whether skeletal features shared by presbyornithids and anhimids are plesiomorphic within Anseriformes or indicative of a close relationship
    between the two cannot yet be resolved. In any case, recognition of W. tedfordi as a presbyornithid calls for a reassessment of the phylogenetic position of Presbyornithidae within Anseriformes."

    Sounds like your hypothesis is much more easily falsifiable than the hypothesis that a
    species of *Merychippus* is directly ancestral to *Equus* in the horse family.

    ??

    Come on, John, the general theme of direct ancestry
    has been a perennial bone of contention between us.
    If you disagree in this instance, please explain your disagreement.

    I truly don't know what you were trying to hint at there, so I don't
    know whether I agree.

    I was not "hinting".

    Still don't know what you were trying to say. What does any of this have
    to do with Merychippus?

    What part of "your hypothesis is much more easily falsifiable than"
    don't you understand? We have essentially complete fossils of Merychippus and can study Equus to our heart's content.

    Compare that to the evidence that *Teviornis gobiensis* was a Cretaceous presbyornithid.
    Mayr's 2017 book doesn't even mention the genus nor the 2016 paper that we have been
    discussing above. Neither does Wikipedia, which only cites two papers on the genus,
    in 2002 and 2004. Do you know of any citations of it in the peer-reviewed literature?
    Or any more recent evidence for its tentative conclusion?

    It's an awfully thin peg to hang 7 separate lineages on.

    Molecular studies calibrated using Cenozoic fossils have more. So it
    seems a conservative number.

    One might also add Gastornithidae and Pelagornithidae, though I'm not >>>>>> sure whether those are both separate branches.

    Ah, one of my favorite bird families, those pelagornithids ["pseudotoothed birds"].
    The biggest one of all, with a wingspread over 20 feet, was found right here
    in South Carolina, while clearing land for the Charleston international airport.

    I'm glad you mentioned them. I found out a fascinating piece of information from the
    Wikipedia entry on them just now.

    "They were the dominant seabirds of most oceans throughout most of the Cenozoic, and modern humans apparently missed encountering them only by a tiny measure of evolutionary time: the last known pelagornithids were contemporaries of Homo habilis and
    the beginning of the history of technology.[3]"

    So near and yet so far. :(

    Similar story for phorusrhacids.

    Even more so: put Homo erectus there. However, the much more recent claims of ca.100 kya
    and even 18 kya are not well supported according to the entry on them.
    The summary in the box at upper right should have a question mark added after 0.1 Ma.
    Another case of discordant statements on the same Wiki page.


    However, I saw no candidates for either family having Cretaceous members. >>>>> And whether they are separate branches or the same seems irrelevant. >>>>> You really have your work cut out for you to find an indirect phylogenetic
    tree way to make them survivors of the K-P extinction.

    You still don't seem to understand how this works.

    I don't understand why you say this. You are ignoring the fact that Gastornis
    is considered by some to be within crown Anseriformes.

    I'm not aware of that. Who?

    Read the Wikipedia entry.

    Or you could just show the citation for that claim. Then I wouldn't have
    to guess which Wikipedia entry you meant.

    As for pelagornithids, the Wikipedia entry tentatively put them into Galloanserae,
    but then treated us to a long discussion that reads like a technical review paper
    of various proposed possibilities, including some "outliers" in Neoaves.

    Sounds like it was written by a very old specialist. Better to look for
    pubs with actual phylogenetic analyses in them, none of which I have
    off-hand.

    Please let us know when you find any. But keep in mind that the
    reviewer cites a number of them already.

    No problem, then.

    If we have a
    Cretaceous example of a particular branch, all nodes below that branch >>>> must also exist at that time. Given a Cretaceous presbyornithid, and
    give tnat Presbyornithidae is the sister group of Anatidae, there must >>>> be Cretaceous gastornithids and/or (depending on the tree)
    pelagornithids, and anseranatids, anhimids, neoavians, and paleognaths. >>>
    Too many assumptions here, in the light of my latest comments.
    Add to them the possibility that Mayr gave [see my first reply to your post],
    that presbyornithids are outside Anseriformes, and you could be down to
    paleognaths, neoaves, galliforms, presbyornithids, and a cladogenesis
    at some Paleocene member of total Anseriformes after anagenesis
    (or evolutionary stasis) from where presbyornithids split off from from Anseriformes.

    Five lineages, not seven.

    Yes, that's possible, though whatever you are trying to say regarding
    anagenesis is unclear.

    One or more speciations without branching. The old species may
    linger on for a while, but in the absence of fossils for side branches,
    there is no evidence for cladogenesis.

    That's not what "speciation" means. Anagenesis is just evolution within
    a lineage. Of course there is a conflict of species concepts here; the morphospecies often used by paleontologists are not species as commonly understood by those who study speciation.

    We're talking about lineages, not individual taxa.

    Not clear on what that was supposed to mean either.

    Just "anseriforms" would have been simpler and
    more correct.

    How so? doesn't it mean the same thing as Anseriformes?

    I'm saying you could have left out all the stuff about "after
    anagenesis, etc.

    So we're agreed that you don't need more than one
    Cretaceous fossil to infer the Cretaceous existence of several other
    groups.

    Of course, but the fossil involved in this case is fragmentary.

    Does that prevent it from being diagnostic?

    I've asked Trevor Worthy last night (my time; this morning his time)
    in email about how the 2016 paper has fared since then, and I hope he answers before long.


    Peter Nyikos
    Professor, Dept. of Mathematics -- standard disclaimer--
    U. of South Carolina at Columbia
    http://people.math.sc.edu/nyikos

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From Peter Nyikos@21:1/5 to daud....@gmail.com on Fri Dec 2 17:53:29 2022
    Short on time, I address both of the first two posts listed below:

    On Friday, December 2, 2022 at 8:08:42 PM UTC-5, daud....@gmail.com wrote:
    On Friday, December 2, 2022 at 7:52:41 PM UTC-5, John Harshman wrote:
    On 12/1/22 10:18 PM, Daud Deden wrote:
    On Wednesday, November 9, 2022 at 9:14:07 PM UTC-5, peter2...@gmail.com wrote:
    The Great K-T extinction spelled the doom of the giant sea reptiles, all dinosaurs
    (unless you count birds as dinosaurs), and many other groups of vertebrates and invertebrates.
    In fact, the damage caused by the asteroid that hit Yucatan was so great
    that no animal over 50 kilos is known to have survived.

    But some groups of birds and mammals did survive, including the remote >> ancestors
    of the four groups that have survived to this day: the neornithine birds and three groups of mammals:
    the placentals, the marsupials, and the monotremes.

    To keep this OP reasonably short, I will deal with the two most familiar of the four, the placentals and the modern birds.

    A 2013 _Science_ article [1] led to the general acceptance of the hypothesis that all living placentals
    have descended from a common ancestor that lived after the great K-T disaster.

    However, there are closely related eutherians [2] that also survived for a while, among
    whose descendants were the pantodonts, including *Coryphodon*, which was a bit larger than a tapir; and the tillodonts and taeniodonts. These groups died out during the Eocene, which ended almost halfway between the K-T disaster and our time.

    The case of birds is generally believed to be quite different: EVERY bird, living
    or extinct, that survived the K-T disaster is believed to be descended from the
    last common ancestor of living birds. However, there are some articles which
    hypothesize that there is one exception, a group of birds called the lithornids.
    More about them in my next post to this thread, some time tomorrow.

    [1] Maureen A. O'Leary et. al., "The Placental Mammal Ancestor and the Post–K-Pg Radiation of Placentals". Science. 339 (6120): 662–667.

    [2] Eutherians are the mammals, extant and extinct, that are more closely related to placentals than to marsupials.
    Similarly, metatherians are the mammals that are more closely related to marsupials than to placentals.


    Peter Nyikos
    Professor, Dept. of Mathematics -- standard disclaimer--
    University of South Carolina
    http://people.math.sc.edu/nyikos

    PS The "Why?" at the end of the thread title is two-edged: why did some survive,
    and why did others, including some similar-seeming ones, perish?
    -
    Not sure where to put this: https://www.nature.com/articles/d41586-022-04181-7

    As usual in science journalism, the title (at least) is hype.

    I don't know why John said this. Chances are he didn't look at the research article
    that Erik linked for us. I skimmed it for less than a minute, and the following
    leaped out at me:

    "Nonetheless, the notably galloanseran-like pterygoid of Janavis, together with recent evidence of a neognath-like hemipterygoid–palatine complex in Ichthyornis12, rejects the hypothesis that the plesiomorphic condition of the neornithine palate was
    palaeognathous. Instead, these observations support the hypothesis that the extant palaeognathous condition is derived and convergent with the superficially similar morphologies found in stemward avialans and non-avialan theropods. Indeed, the pronounced
    morphometric disparity among extant palaeognath pterygoids (Fig. 3 and Extended Data Figs. 8 and 9) further emphasizes that the ‘palaeognathous condition’ may be little more than an anachronistic conceptual wastebasket, oversimplifying the
    substantial morphological variability within this taxonomically depauperate extant clade, and perhaps reflecting outdated assumptions that palaeognaths represent ‘primitive avian stock’ carrying over from the early days of systematic ornithology4."

    --"Cretaceous ornithurine supports a neognathous crown bird ancestor" https://www.nature.com/articles/s41586-022-05445-y


    Why, the very title should have made John think twice before posting the following:

    This discovery has no effect on the evolutionary tree, just on the character
    state at the base of the tree. Paleognaths and neognaths are still each monophyletic. No change there.

    Unfortunately for John, Erik did not post the title, only the nondescriptive url.
    And so we have this priceless irony.


    OT streamlined theropod natovenator no wings https://www.nature.com/articles/s42003-022-04119-9

    It is OT for this thread, but perfectly on-topic for sci.bio.paleontology,
    and another fine catch by you, Daud. Why didn't you begin another thread with it?
    There are so few threads active here, another one is always welcome when the article talked about in the OP is so interesting.


    Peter Nyikos
    Professor, Dept. of Mathematics
    U. of South Carolina -- standard disclaimer--
    http://people.math.sc.edu/nyikos

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From John Harshman@21:1/5 to All on Fri Dec 2 19:29:03 2022
    T24gMTIvMi8yMiA1OjUzIFBNLCBQZXRlciBOeWlrb3Mgd3JvdGU6DQo+IFNob3J0IG9uIHRp bWUsIEkgYWRkcmVzcyBib3RoIG9mIHRoZSBmaXJzdCB0d28gcG9zdHMgbGlzdGVkIGJlbG93 Og0KPiANCj4gT24gRnJpZGF5LCBEZWNlbWJlciAyLCAyMDIyIGF0IDg6MDg6NDIgUE0gVVRD LTUsIGRhdWQuLi4uQGdtYWlsLmNvbSB3cm90ZToNCj4+IE9uIEZyaWRheSwgRGVjZW1iZXIg MiwgMjAyMiBhdCA3OjUyOjQxIFBNIFVUQy01LCBKb2huIEhhcnNobWFuIHdyb3RlOg0KPj4+ IE9uIDEyLzEvMjIgMTA6MTggUE0sIERhdWQgRGVkZW4gd3JvdGU6DQo+Pj4+IE9uIFdlZG5l c2RheSwgTm92ZW1iZXIgOSwgMjAyMiBhdCA5OjE0OjA3IFBNIFVUQy01LCBwZXRlcjIuLi5A Z21haWwuY29tIHdyb3RlOg0KPj4+Pj4gVGhlIEdyZWF0IEstVCBleHRpbmN0aW9uIHNwZWxs ZWQgdGhlIGRvb20gb2YgdGhlIGdpYW50IHNlYSByZXB0aWxlcywgYWxsIGRpbm9zYXVycw0K Pj4+Pj4gKHVubGVzcyB5b3UgY291bnQgYmlyZHMgYXMgZGlub3NhdXJzKSwgYW5kIG1hbnkg b3RoZXIgZ3JvdXBzIG9mIHZlcnRlYnJhdGVzIGFuZCBpbnZlcnRlYnJhdGVzLg0KPj4+Pj4g SW4gZmFjdCwgdGhlIGRhbWFnZSBjYXVzZWQgYnkgdGhlIGFzdGVyb2lkIHRoYXQgaGl0IFl1 Y2F0YW4gd2FzIHNvIGdyZWF0DQo+Pj4+PiB0aGF0IG5vIGFuaW1hbCBvdmVyIDUwIGtpbG9z IGlzIGtub3duIHRvIGhhdmUgc3Vydml2ZWQuDQo+Pj4+Pg0KPj4+Pj4gQnV0IHNvbWUgZ3Jv dXBzIG9mIGJpcmRzIGFuZCBtYW1tYWxzIGRpZCBzdXJ2aXZlLCBpbmNsdWRpbmcgdGhlIHJl bW90ZQ0KPj4+Pj4gYW5jZXN0b3JzDQo+Pj4+PiBvZiB0aGUgZm91ciBncm91cHMgdGhhdCBo YXZlIHN1cnZpdmVkIHRvIHRoaXMgZGF5OiB0aGUgbmVvcm5pdGhpbmUgYmlyZHMgYW5kIHRo cmVlIGdyb3VwcyBvZiBtYW1tYWxzOg0KPj4+Pj4gdGhlIHBsYWNlbnRhbHMsIHRoZSBtYXJz dXBpYWxzLCBhbmQgdGhlIG1vbm90cmVtZXMuDQo+Pj4+Pg0KPj4+Pj4gVG8ga2VlcCB0aGlz IE9QIHJlYXNvbmFibHkgc2hvcnQsIEkgd2lsbCBkZWFsIHdpdGggdGhlIHR3byBtb3N0IGZh bWlsaWFyIG9mIHRoZSBmb3VyLCB0aGUgcGxhY2VudGFscyBhbmQgdGhlIG1vZGVybiBiaXJk cy4NCj4+Pj4+DQo+Pj4+PiBBIDIwMTMgX1NjaWVuY2VfIGFydGljbGUgWzFdIGxlZCB0byB0 aGUgZ2VuZXJhbCBhY2NlcHRhbmNlIG9mIHRoZSBoeXBvdGhlc2lzIHRoYXQgYWxsIGxpdmlu ZyBwbGFjZW50YWxzDQo+Pj4+PiBoYXZlIGRlc2NlbmRlZCBmcm9tIGEgY29tbW9uIGFuY2Vz dG9yIHRoYXQgbGl2ZWQgYWZ0ZXIgdGhlIGdyZWF0IEstVCBkaXNhc3Rlci4NCj4+Pj4+DQo+ Pj4+PiBIb3dldmVyLCB0aGVyZSBhcmUgY2xvc2VseSByZWxhdGVkIGV1dGhlcmlhbnMgWzJd IHRoYXQgYWxzbyBzdXJ2aXZlZCBmb3IgYSB3aGlsZSwgYW1vbmcNCj4+Pj4+IHdob3NlIGRl c2NlbmRhbnRzIHdlcmUgdGhlIHBhbnRvZG9udHMsIGluY2x1ZGluZyAqQ29yeXBob2Rvbios IHdoaWNoIHdhcyBhIGJpdCBsYXJnZXIgdGhhbiBhIHRhcGlyOyBhbmQgdGhlIHRpbGxvZG9u dHMgYW5kIHRhZW5pb2RvbnRzLiBUaGVzZSBncm91cHMgZGllZCBvdXQgZHVyaW5nIHRoZSBF b2NlbmUsIHdoaWNoIGVuZGVkIGFsbW9zdCBoYWxmd2F5IGJldHdlZW4gdGhlIEstVCBkaXNh c3RlciBhbmQgb3VyIHRpbWUuDQo+Pj4+Pg0KPj4+Pj4gVGhlIGNhc2Ugb2YgYmlyZHMgaXMg Z2VuZXJhbGx5IGJlbGlldmVkIHRvIGJlIHF1aXRlIGRpZmZlcmVudDogRVZFUlkgYmlyZCwg bGl2aW5nDQo+Pj4+PiBvciBleHRpbmN0LCB0aGF0IHN1cnZpdmVkIHRoZSBLLVQgZGlzYXN0 ZXIgaXMgYmVsaWV2ZWQgdG8gYmUgZGVzY2VuZGVkIGZyb20gdGhlDQo+Pj4+PiBsYXN0IGNv bW1vbiBhbmNlc3RvciBvZiBsaXZpbmcgYmlyZHMuIEhvd2V2ZXIsIHRoZXJlIGFyZSBzb21l IGFydGljbGVzIHdoaWNoDQo+Pj4+PiBoeXBvdGhlc2l6ZSB0aGF0IHRoZXJlIGlzIG9uZSBl eGNlcHRpb24sIGEgZ3JvdXAgb2YgYmlyZHMgY2FsbGVkIHRoZSBsaXRob3JuaWRzLg0KPj4+ Pj4gTW9yZSBhYm91dCB0aGVtIGluIG15IG5leHQgcG9zdCB0byB0aGlzIHRocmVhZCwgc29t ZSB0aW1lIHRvbW9ycm93Lg0KPj4+Pj4NCj4+Pj4+IFsxXSBNYXVyZWVuIEEuIE8nTGVhcnkg ZXQuIGFsLiwgIlRoZSBQbGFjZW50YWwgTWFtbWFsIEFuY2VzdG9yIGFuZCB0aGUgUG9zdOKA k0stUGcgUmFkaWF0aW9uIG9mIFBsYWNlbnRhbHMiLiBTY2llbmNlLiAzMzkgKDYxMjApOiA2 NjLigJM2NjcuDQo+Pj4+Pg0KPj4+Pj4gWzJdIEV1dGhlcmlhbnMgYXJlIHRoZSBtYW1tYWxz LCBleHRhbnQgYW5kIGV4dGluY3QsIHRoYXQgYXJlIG1vcmUgY2xvc2VseSByZWxhdGVkIHRv IHBsYWNlbnRhbHMgdGhhbiB0byBtYXJzdXBpYWxzLg0KPj4+Pj4gU2ltaWxhcmx5LCBtZXRh dGhlcmlhbnMgYXJlIHRoZSBtYW1tYWxzIHRoYXQgYXJlIG1vcmUgY2xvc2VseSByZWxhdGVk IHRvIG1hcnN1cGlhbHMgdGhhbiB0byBwbGFjZW50YWxzLg0KPj4+Pj4NCj4+Pj4+DQo+Pj4+ PiBQZXRlciBOeWlrb3MNCj4+Pj4+IFByb2Zlc3NvciwgRGVwdC4gb2YgTWF0aGVtYXRpY3Mg LS0gc3RhbmRhcmQgZGlzY2xhaW1lci0tDQo+Pj4+PiBVbml2ZXJzaXR5IG9mIFNvdXRoIENh cm9saW5hDQo+Pj4+PiBodHRwOi8vcGVvcGxlLm1hdGguc2MuZWR1L255aWtvcw0KPj4+Pj4N Cj4+Pj4+IFBTIFRoZSAiV2h5PyIgYXQgdGhlIGVuZCBvZiB0aGUgdGhyZWFkIHRpdGxlIGlz IHR3by1lZGdlZDogd2h5IGRpZCBzb21lIHN1cnZpdmUsDQo+Pj4+PiBhbmQgd2h5IGRpZCBv dGhlcnMsIGluY2x1ZGluZyBzb21lIHNpbWlsYXItc2VlbWluZyBvbmVzLCBwZXJpc2g/DQo+ Pj4+IC0NCj4+Pj4gTm90IHN1cmUgd2hlcmUgdG8gcHV0IHRoaXM6IGh0dHBzOi8vd3d3Lm5h dHVyZS5jb20vYXJ0aWNsZXMvZDQxNTg2LTAyMi0wNDE4MS03DQo+IA0KPj4+IEFzIHVzdWFs IGluIHNjaWVuY2Ugam91cm5hbGlzbSwgdGhlIHRpdGxlIChhdCBsZWFzdCkgaXMgaHlwZS4N Cj4gDQo+IEkgZG9uJ3Qga25vdyB3aHkgSm9obiBzYWlkIHRoaXMuDQoNCkFuZCBJIGRvbid0 IGtub3cgd2h5IFBldGVyIGNob29zZXMgdG8gcmVwbHkgYXQgc2Vjb25kLWhhbmQgcmF0aGVy IHRoYW4gDQp0byB0aGUgYWN0dWFsIHBvc3QgaGUncyByZXBseWluZyB0by4gSWYgb25seSBo ZSB3b3VsZCBzdG9wLg0KDQo+IENoYW5jZXMgYXJlIGhlIGRpZG4ndCBsb29rIGF0IHRoZSBy ZXNlYXJjaCBhcnRpY2xlDQo+IHRoYXQgRXJpayBsaW5rZWQgZm9yIHVzLiBJIHNraW1tZWQg aXQgZm9yIGxlc3MgdGhhbiBhIG1pbnV0ZSwgYW5kIHRoZSBmb2xsb3dpbmcNCj4gbGVhcGVk IG91dCBhdCBtZToNCj4gDQo+ICJOb25ldGhlbGVzcywgdGhlIG5vdGFibHkgZ2FsbG9hbnNl cmFuLWxpa2UgcHRlcnlnb2lkIG9mIEphbmF2aXMsIHRvZ2V0aGVyIHdpdGggcmVjZW50IGV2 aWRlbmNlIG9mIGEgbmVvZ25hdGgtbGlrZSBoZW1pcHRlcnlnb2lk4oCTcGFsYXRpbmUgY29t cGxleCBpbiBJY2h0aHlvcm5pczEyLCByZWplY3RzIHRoZSBoeXBvdGhlc2lzIHRoYXQgdGhl IHBsZXNpb21vcnBoaWMgY29uZGl0aW9uIG9mIHRoZSBuZW9ybml0aGluZSBwYWxhdGUgd2Fz IHBhbGFlb2duYXRob3VzLiBJbnN0ZWFkLCB0aGVzZSBvYnNlcnZhdGlvbnMgc3VwcG9ydCB0 aGUgaHlwb3RoZXNpcyB0aGF0IHRoZSBleHRhbnQgcGFsYWVvZ25hdGhvdXMgY29uZGl0aW9u IGlzIGRlcml2ZWQgYW5kIGNvbnZlcmdlbnQgd2l0aCB0aGUgc3VwZXJmaWNpYWxseSBzaW1p bGFyIG1vcnBob2xvZ2llcyBmb3VuZCBpbiBzdGVtd2FyZCBhdmlhbGFucyBhbmQgbm9uLWF2 aWFsYW4gdGhlcm9wb2RzLiBJbmRlZWQsIHRoZSBwcm9ub3VuY2VkIG1vcnBob21ldHJpYyBk aXNwYXJpdHkgYW1vbmcgZXh0YW50IHBhbGFlb2duYXRoIHB0ZXJ5Z29pZHMgKEZpZy4gMyBh bmQgRXh0ZW5kZWQgRGF0YSBGaWdzLiA4IGFuZCA5KSBmdXJ0aGVyIGVtcGhhc2l6ZXMgdGhh dCB0aGUg4oCYcGFsYWVvZ25hdGhvdXMgY29uZGl0aW9u4oCZIG1heSBiZSBsaXR0bGUgbW9y ZSB0aGFuIGFuIGFuYWNocm9uaXN0aWMgY29uY2VwdHVhbCB3YXN0ZWJhc2tldCwgb3ZlcnNp bXBsaWZ5aW5nIHRoZSBzdWJzdGFudGlhbCBtb3JwaG9sb2dpY2FsIHZhcmlhYmlsaXR5IHdp dGhpbiB0aGlzIHRheG9ub21pY2FsbHkgZGVwYXVwZXJhdGUgZXh0YW50IGNsYWRlLCBhbmQg cGVyaGFwcyByZWZsZWN0aW5nIG91dGRhdGVkIGFzc3VtcHRpb25zIHRoYXQgcGFsYWVvZ25h dGhzIHJlcHJlc2VudCDigJhwcmltaXRpdmUgYXZpYW4gc3RvY2vigJkgY2Fycnlpbmcgb3Zl ciBmcm9tIHRoZSBlYXJseSBkYXlzIG9mIHN5c3RlbWF0aWMgb3JuaXRob2xvZ3k0LiINCj4g DQo+IC0tIkNyZXRhY2VvdXMgb3JuaXRodXJpbmUgc3VwcG9ydHMgYSBuZW9nbmF0aG91cyBj cm93biBiaXJkIGFuY2VzdG9yIg0KPiBodHRwczovL3d3dy5uYXR1cmUuY29tL2FydGljbGVz L3M0MTU4Ni0wMjItMDU0NDUteQ0KPiANCj4gDQo+IFdoeSwgdGhlIHZlcnkgdGl0bGUgc2hv dWxkIGhhdmUgbWFkZSBKb2huIHRoaW5rIHR3aWNlIGJlZm9yZSAgcG9zdGluZyB0aGUgZm9s bG93aW5nOg0KPiANCj4+PiBUaGlzIGRpc2NvdmVyeSBoYXMgbm8gZWZmZWN0IG9uIHRoZSBl dm9sdXRpb25hcnkgdHJlZSwganVzdCBvbiB0aGUgY2hhcmFjdGVyDQo+Pj4gc3RhdGUgYXQg dGhlIGJhc2Ugb2YgdGhlIHRyZWUuIFBhbGVvZ25hdGhzIGFuZCBuZW9nbmF0aHMgYXJlIHN0 aWxsIGVhY2gNCj4+PiBtb25vcGh5bGV0aWMuIE5vIGNoYW5nZSB0aGVyZS4NCj4gDQo+IFVu Zm9ydHVuYXRlbHkgZm9yIEpvaG4sIEVyaWsgZGlkIG5vdCBwb3N0IHRoZSB0aXRsZSwgb25s eSAgdGhlIG5vbmRlc2NyaXB0aXZlIHVybC4NCj4gQW5kIHNvIHdlIGhhdmUgdGhpcyBwcmlj ZWxlc3MgaXJvbnkuDQoNCkknbSBzdGlsbCB1bmF3YXJlIG9mIHRoZSBpcm9ueSwgd2hpY2gg eW91IHByb2JhYmx5IGJlbGlldmUsIHdyb25nbHksIHRvIA0KYmUgc2VsZi1ldmlkZW50LiBJ J20gbGVmdCB0byBndWVzcyB0aGF0IHlvdSBtaXN0YWtlIGh5cG90aGVzZXMgb2YgDQpjaGFy YWN0ZXIgZXZvbHV0aW9uIG9uIHRoZSB0cmVlIGZvciB0aGUgdHJlZSBpdHNlbGYuIEFnYWlu LCBldmVuIGlmIHdlIA0KZmluZCB0aGF0IGEgY2hhcmFjdGVyIGF0IHRoZSBiYXNlIG9mIE5l b3JuaXRoZXMgKEF2ZXMpIGlzIG90aGVyIHRoYW4gDQpwcmV2aW91c2x5IHRob3VnaHQsIHRo YXQgZG9lc24ndCBuZWNlc3NhcmlseSBjaGFuZ2UgdGhlIHJlbGF0aW9uc2hpcHMgDQphbW9u ZyB0YXhhLCBhbmQgaXQgbW9zdCBjZXJ0YWlubHkgZG9lcyBub3QgaW4gdGhlIHByZXNlbnQg Y2FzZS4NCg0KIlByaW1pdGl2ZSBhdmlhbiBzdG9jayIgaXMgZGVmaW5pdGVseSBhbiBhbmFj aHJvbmlzdGljIGFuZCBvdXRkYXRlZCANCmNvbmNlcHQuIEkgc3VzcGVjdCB0aGF0IHlvdSBk b24ndCB1bmRlcnN0YW5kIHdoYXQgdGhlIGF1dGhvcnMgd2VyZSANCnNheWluZyB0aGVyZS4N Cg0KDQo=

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From erik simpson@21:1/5 to peter2...@gmail.com on Sat Dec 3 10:20:13 2022
    On Friday, December 2, 2022 at 5:53:31 PM UTC-8, peter2...@gmail.com wrote:
    Short on time, I address both of the first two posts listed below:
    On Friday, December 2, 2022 at 8:08:42 PM UTC-5, daud....@gmail.com wrote:
    On Friday, December 2, 2022 at 7:52:41 PM UTC-5, John Harshman wrote:
    On 12/1/22 10:18 PM, Daud Deden wrote:
    On Wednesday, November 9, 2022 at 9:14:07 PM UTC-5, peter2...@gmail.com wrote:
    The Great K-T extinction spelled the doom of the giant sea reptiles, all dinosaurs
    (unless you count birds as dinosaurs), and many other groups of vertebrates and invertebrates.
    In fact, the damage caused by the asteroid that hit Yucatan was so great
    that no animal over 50 kilos is known to have survived.

    But some groups of birds and mammals did survive, including the remote
    ancestors
    of the four groups that have survived to this day: the neornithine birds and three groups of mammals:
    the placentals, the marsupials, and the monotremes.

    To keep this OP reasonably short, I will deal with the two most familiar of the four, the placentals and the modern birds.

    A 2013 _Science_ article [1] led to the general acceptance of the hypothesis that all living placentals
    have descended from a common ancestor that lived after the great K-T disaster.

    However, there are closely related eutherians [2] that also survived for a while, among
    whose descendants were the pantodonts, including *Coryphodon*, which was a bit larger than a tapir; and the tillodonts and taeniodonts. These groups died out during the Eocene, which ended almost halfway between the K-T disaster and our time.

    The case of birds is generally believed to be quite different: EVERY bird, living
    or extinct, that survived the K-T disaster is believed to be descended from the
    last common ancestor of living birds. However, there are some articles which
    hypothesize that there is one exception, a group of birds called the lithornids.
    More about them in my next post to this thread, some time tomorrow. >>
    [1] Maureen A. O'Leary et. al., "The Placental Mammal Ancestor and the Post–K-Pg Radiation of Placentals". Science. 339 (6120): 662–667.

    [2] Eutherians are the mammals, extant and extinct, that are more closely related to placentals than to marsupials.
    Similarly, metatherians are the mammals that are more closely related to marsupials than to placentals.


    Peter Nyikos
    Professor, Dept. of Mathematics -- standard disclaimer--
    University of South Carolina
    http://people.math.sc.edu/nyikos

    PS The "Why?" at the end of the thread title is two-edged: why did some survive,
    and why did others, including some similar-seeming ones, perish?
    -
    Not sure where to put this: https://www.nature.com/articles/d41586-022-04181-7

    As usual in science journalism, the title (at least) is hype.
    I don't know why John said this. Chances are he didn't look at the research article
    that Erik linked for us. I skimmed it for less than a minute, and the following
    leaped out at me:

    "Nonetheless, the notably galloanseran-like pterygoid of Janavis, together with recent evidence of a neognath-like hemipterygoid–palatine complex in Ichthyornis12, rejects the hypothesis that the plesiomorphic condition of the neornithine palate was
    palaeognathous. Instead, these observations support the hypothesis that the extant palaeognathous condition is derived and convergent with the superficially similar morphologies found in stemward avialans and non-avialan theropods. Indeed, the pronounced
    morphometric disparity among extant palaeognath pterygoids (Fig. 3 and Extended Data Figs. 8 and 9) further emphasizes that the ‘palaeognathous condition’ may be little more than an anachronistic conceptual wastebasket, oversimplifying the
    substantial morphological variability within this taxonomically depauperate extant clade, and perhaps reflecting outdated assumptions that palaeognaths represent ‘primitive avian stock’ carrying over from the early days of systematic ornithology4."

    --"Cretaceous ornithurine supports a neognathous crown bird ancestor" https://www.nature.com/articles/s41586-022-05445-y


    Why, the very title should have made John think twice before posting the following:
    This discovery has no effect on the evolutionary tree, just on the character
    state at the base of the tree. Paleognaths and neognaths are still each monophyletic. No change there.
    Unfortunately for John, Erik did not post the title, only the nondescriptive url.
    And so we have this priceless irony.

    What priceless irony? Maybe you should try reading the article for a little more than
    a minute, and maybe even think a bit about what the "message" is. Hint: consider
    what the term ‘palaeognathous condition’ is, and what its phylogentic implications are.

    The reference given for the "early days of systematic ornithology" is itself a glimpse of
    good old-time phylogeny circa 1948, with references going back as far as Huxley
    himself (1867). Thirty pages of exquisite anaysis of the bony palate of paleognaths.

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)
  • From erik simpson@21:1/5 to erik simpson on Sat Dec 3 10:22:54 2022
    On Saturday, December 3, 2022 at 10:20:14 AM UTC-8, erik simpson wrote:
    On Friday, December 2, 2022 at 5:53:31 PM UTC-8, peter2...@gmail.com wrote:
    Short on time, I address both of the first two posts listed below:
    On Friday, December 2, 2022 at 8:08:42 PM UTC-5, daud....@gmail.com wrote:
    On Friday, December 2, 2022 at 7:52:41 PM UTC-5, John Harshman wrote:
    On 12/1/22 10:18 PM, Daud Deden wrote:
    On Wednesday, November 9, 2022 at 9:14:07 PM UTC-5, peter2...@gmail.com wrote:
    The Great K-T extinction spelled the doom of the giant sea reptiles, all dinosaurs
    (unless you count birds as dinosaurs), and many other groups of vertebrates and invertebrates.
    In fact, the damage caused by the asteroid that hit Yucatan was so great
    that no animal over 50 kilos is known to have survived.

    But some groups of birds and mammals did survive, including the remote
    ancestors
    of the four groups that have survived to this day: the neornithine birds and three groups of mammals:
    the placentals, the marsupials, and the monotremes.

    To keep this OP reasonably short, I will deal with the two most familiar of the four, the placentals and the modern birds.

    A 2013 _Science_ article [1] led to the general acceptance of the hypothesis that all living placentals
    have descended from a common ancestor that lived after the great K-T disaster.

    However, there are closely related eutherians [2] that also survived for a while, among
    whose descendants were the pantodonts, including *Coryphodon*, which was a bit larger than a tapir; and the tillodonts and taeniodonts. These groups died out during the Eocene, which ended almost halfway between the K-T disaster and our time.

    The case of birds is generally believed to be quite different: EVERY bird, living
    or extinct, that survived the K-T disaster is believed to be descended from the
    last common ancestor of living birds. However, there are some articles which
    hypothesize that there is one exception, a group of birds called the lithornids.
    More about them in my next post to this thread, some time tomorrow. >>
    [1] Maureen A. O'Leary et. al., "The Placental Mammal Ancestor and the Post–K-Pg Radiation of Placentals". Science. 339 (6120): 662–667.

    [2] Eutherians are the mammals, extant and extinct, that are more closely related to placentals than to marsupials.
    Similarly, metatherians are the mammals that are more closely related to marsupials than to placentals.


    Peter Nyikos
    Professor, Dept. of Mathematics -- standard disclaimer--
    University of South Carolina
    http://people.math.sc.edu/nyikos

    PS The "Why?" at the end of the thread title is two-edged: why did some survive,
    and why did others, including some similar-seeming ones, perish?
    -
    Not sure where to put this: https://www.nature.com/articles/d41586-022-04181-7

    As usual in science journalism, the title (at least) is hype.
    I don't know why John said this. Chances are he didn't look at the research article
    that Erik linked for us. I skimmed it for less than a minute, and the following
    leaped out at me:

    "Nonetheless, the notably galloanseran-like pterygoid of Janavis, together with recent evidence of a neognath-like hemipterygoid–palatine complex in Ichthyornis12, rejects the hypothesis that the plesiomorphic condition of the neornithine palate
    was palaeognathous. Instead, these observations support the hypothesis that the extant palaeognathous condition is derived and convergent with the superficially similar morphologies found in stemward avialans and non-avialan theropods. Indeed, the
    pronounced morphometric disparity among extant palaeognath pterygoids (Fig. 3 and Extended Data Figs. 8 and 9) further emphasizes that the ‘palaeognathous condition’ may be little more than an anachronistic conceptual wastebasket, oversimplifying the
    substantial morphological variability within this taxonomically depauperate extant clade, and perhaps reflecting outdated assumptions that palaeognaths represent ‘primitive avian stock’ carrying over from the early days of systematic ornithology4."

    --"Cretaceous ornithurine supports a neognathous crown bird ancestor" https://www.nature.com/articles/s41586-022-05445-y


    Why, the very title should have made John think twice before posting the following:
    This discovery has no effect on the evolutionary tree, just on the character
    state at the base of the tree. Paleognaths and neognaths are still each
    monophyletic. No change there.
    Unfortunately for John, Erik did not post the title, only the nondescriptive url.
    And so we have this priceless irony.
    What priceless irony? Maybe you should try reading the article for a little more than
    a minute, and maybe even think a bit about what the "message" is. Hint: consider
    what the term ‘palaeognathous condition’ is, and what its phylogentic implications are.

    The reference given for the "early days of systematic ornithology" is itself a glimpse of
    good old-time phylogeny circa 1948, with references going back as far as Huxley
    himself (1867). Thirty pages of exquisite anaysis of the bony palate of paleognaths.

    Forgot the reference itself: https://academic.oup.com/auk/article/65/4/520/5247317

    --- SoupGate-Win32 v1.05
    * Origin: fsxNet Usenet Gateway (21:1/5)