• Lucy was aquarboreal

    From littoral.homo@gmail.com@21:1/5 to All on Tue Dec 13 04:24:33 2022
    Morphological correlates of distal fibular morphology with locomotion in great apes, humans, and Australopithecus afarensis
    Damiano Marchi cs 2022 Am.J.biol.Anthr. doi org/10.1002/ajpa.24507
    "... Au.afarensis shows a unique distal fibular morphology, with several traits generally associated more to arboreality, less to BPism. ..."

    arboreal + bipedal = aquarboreal

    Lucy was no human ancestor, of course, but a Pliocene relative of gorillas: Gorilla fossil subspecies Praeanthropus afarensis

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  • From JTEM is so reasonable@21:1/5 to littor...@gmail.com on Tue Dec 13 14:21:22 2022
    littor...@gmail.com wrote:

    arboreal + bipedal = aquarboreal

    Lucy was no human ancestor, of course, but a Pliocene relative of gorillas: Gorilla fossil subspecies Praeanthropus afarensis

    According to your model -- and my model! -- Lucy descended from an
    earlier Waterside ancestor. So we'd expect aquatic adaptations. We
    could help settle frayed nerves in the savanna cult by saying she was "Transitional," but that's not entirely correct.

    I despise linear models.

    Whatever Lucy's lifestyle, her anatomy seemed to fit it just fine. Her
    species is credited with surviving more than twice as long as the
    savanna cult claims for so called "modern" humans. And that included
    the retention of aquatic traits? So what. As I've pointed out many
    times; giant sauropod dinosaurs retained skeletal features of a much
    smaller, bipedal dinosaur. Unless there's both selective pressure
    AGAINST adaptation, AND the genetic capacity (diversity? potential?),
    things are going to stay exactly the same.

    So I wouldn't be quick to claim she was aquarboreal. Even if she wasn't,
    having descended from Waterside ancestors we couldn't expect her to
    look much if any different.




    -- --

    https://jtem.tumblr.com/post/703468006592905216

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  • From littoral.homo@gmail.com@21:1/5 to All on Tue Dec 13 15:48:51 2022
    Op dinsdag 13 december 2022 om 23:21:23 UTC+1 schreef JTEM is so reasonable:
    littor...@gmail.com wrote:

    arboreal + bipedal = aquarboreal
    Lucy was no human ancestor, of course, but a Pliocene relative of gorillas:
    Gorilla fossil subspecies Praeanthropus afarensis

    Sorry, I meant, of course, "fossil subgenus".

    According to your model -- and my model! -- Lucy descended from an
    earlier Waterside ancestor. So we'd expect aquatic adaptations. We
    could help settle frayed nerves in the savanna cult by saying she was "Transitional," but that's not entirely correct.

    There's 0 doubt (except to savanna fools) that human ancestors were a lot more aquatic than we are today,
    but since primates are arboreal, and since evolution is in small steps, there had to be something between, which spent time both in the trees & in the water.
    I had proposed this early 1990s IIRC (the term "aquarboreal" is not mine, and came several years later IIRC),
    and indeed (c 1995 or so?), the wading lowland gorillas of Ndoki were discovered, and a few years later also the wading bonobos & the wading orangutans.

    The idea that apiths were Afr.ape ancestors or relatives was not mine (e.g. Oxnard), and initially I didn't believe it
    (most of those "heretics" thought gracile apiths were relatives of Pan, and robust apiths, of Gorilla),
    but I read lots & lots of PA articles & books, and it soon became clear:
    - most if not all apiths are not found in dry savanna, but in wet forests etc., - E.Afr.apiths (gracile & robust) looked most like gorillas,
    - S.Afr.apiths (gracile & robust) most like chimps or bonobos,
    - IOW, boisei & robustus are not very close relatives: they became "robust" in //.
    And their humanlike traits ("bipedality" etc.) were primitive, not derived. Traditional PAs assume (because of these humanlike traits) that apiths are closer (but extinct) relatives of us than of Afr.apes, and call them "hominin".
    The term "hominin" is usu. or even always superfluous & confusing & prepossessed, e.g. most articles that use "hominin" are already prepossessed.

    From my 1990 Hum.Evol.paper already:
    • The australopith dentition is more apelike in development pattern (Conroy & Vannier 1987), enamel growth rate (Bromage & Dean 1985), dental morphology (Johanson & Edey 1981) and enamel microwear.
    • All australopithecine brain endocasts appear to be ape- rather than humanlike in size and sulcal pattern (Falk 1985).
    • The composite A.afarensis skull (mostly AL 333; Kimbel et al.1984) «looked very much like a small female gorilla» (Johanson & Edey 1981).
    • The extensive pneumatization of the AL 333-45 temporal bone is also seen in chimpanzee males and some gorillas; «the pattern of pneumatization in A. afarensis is also found only in the extant apes among other hominoids» (Kimbel et al.1984).
    • KNM-WT 17000 had «extremely convex inferolateral margins of the orbits such as found in some gorillas» (Walker et al.1986).
    • «The ‘keystone’ nasal bone arrangement suggested as a derived pattern diagnostic of Paranthropus is found in an appreciable number of pongids, particularly clearly in some chimpanzees» (Eckhardt, 1987).
    • A.robustus incus SK 848 resembles Pan more than Homo and certainly than Gorilla (Fig. 1 in Rak & Clarke, 1979).
    • A.africanus Sts 5 resembles a bonobo skull (Zihlman et al.1978).
    • The Taung skull has much more chimp than human traits (Bromage 1985) and is indeed too recent (Partridge 1985) to be on the line to Homo.
    • Its «pneumatization has also extended into the zygoma and hard palate. This is intriguing because an intrapalatal extension of the maxillary sinus has only been reported in chimpanzees and robust australopithecines among higher primates» (Conroy &
    Vannier 1987).



    I despise linear models.
    Whatever Lucy's lifestyle, her anatomy seemed to fit it just fine. Her species is credited with surviving more than twice as long as the
    savanna cult claims for so called "modern" humans. And that included
    the retention of aquatic traits? So what. As I've pointed out many
    times; giant sauropod dinosaurs retained skeletal features of a much smaller, bipedal dinosaur. Unless there's both selective pressure
    AGAINST adaptation, AND the genetic capacity (diversity? potential?),
    things are going to stay exactly the same.
    So I wouldn't be quick to claim she was aquarboreal. Even if she wasn't, having descended from Waterside ancestors we couldn't expect her to
    look much if any different.
    https://jtem.tumblr.com/post/703468006592905216

    Our most-aquatic phase (H.erectus AFAWK) was probably early-Pleistocene.

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  • From JTEM is so reasonable@21:1/5 to littor...@gmail.com on Tue Dec 13 16:46:42 2022
    littor...@gmail.com wrote:

    There's 0 doubt (except to savanna fools) that human ancestors were a lot more aquatic than we are today,
    but since primates are arboreal, and since evolution is in small steps, there had to be something between,
    which spent time both in the trees & in the water.

    All very true, but if you look at the dating for even some of the oldest
    points of divergence, the LCA was Waterside. Chimps became
    arboreal AFTER divergence. Not before. Chimps had to adapt to the
    forest, the trees only AFTER divergence. So if Chimps evolved from
    something that looked like Lucy, Lucy had ancestors who were NOT
    arboreal at all. Lucy is an Aquatic Ape, so to speak, that pushed
    inland and adapted to a new environment... possibly with the help
    of interbreeding with earlier arrivals.

    We know this model is correct, because we do have Neanderthals
    (Denisovans and all the others) so it's just a matter of extrapolating
    what we already know to a previous point along the timeline.

    The idea that apiths were Afr.ape ancestors or relatives was not mine

    I'm not disputing it. I actually don't have a horse in that race, whether
    it was Lucy or something else very close to Lucy the model doesn't
    change.

    We have a Waterside population spreading across the globe, with
    groups periodically pushing inland where they stay and adapt.. resulting
    in diversification. It's also why the Rift Valley is exactly the kind of
    place where we should find fossils! It had an outlet to the sea. It's
    where a Waterside group would push inland, if one were there and
    inclined to do so. All we need for this scenario is conflict, or disease,
    a natural disaster, really bad weather -- SOMETHING that makes them
    "Turn Right" and follow that water inland rather than stay put on the
    coast.

    That's it.

    Of course this would have all been turbocharged once the
    glacial/interglacial cycle was in full swing. We'd have roughly.. I dunno. Maybe 70,000 years of open highways (the continental shelf when
    sea levels dropped) punctuated by maybe 10,000 years of isolation, as
    the interglacial hit, raising sea levels and cutting everyone off from
    each other.

    The key here is Aquatic Ape.

    The most important piece, the one population that is common to all,
    that is everyone's ancestor is that Waterside population! They were
    the conduit. As they encountered these far flung groups, interbreeding, exchanging DNA (acquiring advantageous adaptations) they carried
    them along to the next group they encountered... and the next...

    We only say "Out of Africa" because of Toba. If Yellowstone explodes
    tomorrow it'll be like 50 Krakatoas exploding all at once. When Toba
    erupted it was roughly 2.8x Yellowstone!

    https://jtem.tumblr.com/image/176437002088

    (Not my image. I stole it off the web. But anyone can Google it if they
    doubt what I'm saying)

    THIS is why we say "Out of Africa."

    Toba exploded, Ground Zero was Sundaland -- the "Asia" in the "Out of
    Asia" scenario -- so the other major population center situated along
    the equator, away from where the worst of it was going to be, was
    Africa.

    There. That's the model. That's the fusion of Multiregionalism/Regional Continuity, Punctuated Equilibrium, Coastal Dispersal: Aquatic Ape. Lucy
    is an expected (predicted) biproduct of all that. She descended from a
    group that peeled off of an earlier group, pushed inland and adapted to
    a new environment... resulting in speciation.

    Our most-aquatic phase (H.erectus AFAWK) was probably early-Pleistocene.

    I think you're too conservative. I'd say that erectus is it. "She sang."




    -- --

    https://jtem.tumblr.com/post/176437002088

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  • From littoral.homo@gmail.com@21:1/5 to All on Wed Dec 14 06:16:20 2022
    Op woensdag 14 december 2022 om 01:46:43 UTC+1 schreef JTEM is so reasonable:
    littor...@gmail.com wrote:

    There's 0 doubt (except to savanna fools) that human ancestors were a lot more aquatic than we are today,
    but since primates are arboreal, and since evolution is in small steps, there had to be something between,
    which spent time both in the trees & in the water.

    All very true, but if you look at the dating for even some of the oldest points of divergence, the LCA was Waterside. Chimps became
    arboreal AFTER divergence. Not before. Chimps had to adapt to the
    forest, the trees only AFTER divergence. So if Chimps evolved from
    something that looked like Lucy, Lucy had ancestors who were NOT
    arboreal at all. Lucy is an Aquatic Ape, so to speak, that pushed
    inland and adapted to a new environment... possibly with the help
    of interbreeding with earlier arrivals.

    1) Lucy was at most only a partially "aquatic" ape, more precisely, it was an *aquarboreal* ape,
    something like (google) "wading gorilla". Ape ancestors have alway remained partly arboreal.
    IMO, Miocene Hominoidea spread aquarboreally along Tethys Ocean (later Ind.Ocean + Med.Sea).
    2) The more aquatic (shallow-diving) Homo evolved later, late-Pliocene or possibly only early-Pleist.:
    Pleist.Homo reached overseas islands, evolved pachyosteosclerosis, large brains, platypelloidy etc.etc.

    We know this model is correct, because we do have Neanderthals
    (Denisovans and all the others) so it's just a matter of extrapolating
    what we already know to a previous point along the timeline.

    The idea that apiths were Afr.ape ancestors or relatives was not mine

    I'm not disputing it. I actually don't have a horse in that race, whether
    it was Lucy or something else very close to Lucy the model doesn't
    change.
    We have a Waterside population spreading across the globe, with
    groups periodically pushing inland where they stay and adapt.. resulting
    in diversification. It's also why the Rift Valley is exactly the kind of place where we should find fossils! It had an outlet to the sea. It's
    where a Waterside group would push inland, if one were there and
    inclined to do so. All we need for this scenario is conflict, or disease,
    a natural disaster, really bad weather -- SOMETHING that makes them
    "Turn Right" and follow that water inland rather than stay put on the
    coast.

    Yes, Gorilla & HP split c 8 Ma, when the northern part of the E.Afr.Rift formed (Red Sea),
    Lucy's ancestors followed the Rift ->Gorilla.
    Much later, Taung's ancestors followed the southern part of the E.Afr.Rift ->Pan.
    Gorilla & Pan evolved in //
    - late-Pliocene "gracile" afarensis//africanus,
    - early-Pleist. "robust" boisei//robustus,
    - today Afr.apes gorillas//chimps-bonobos.

    I'd think Homo & Pan split when the Red Sea opened into the Ind.Ocean:
    -- Pan went right = E.Afr.coastal forests initially,
    -- Homo went left = S.Asian coastal forests initially ->shallow-diving early-Pleist.H.erectus Java etc.

    *If* there ever was an "Out-of-Africa", it was late-Pleistocene, and perhaps not even all H.sapiens.

    Of course this would have all been turbocharged once the
    glacial/interglacial cycle was in full swing. We'd have roughly.. I dunno. Maybe 70,000 years of open highways (the continental shelf when
    sea levels dropped) punctuated by maybe 10,000 years of isolation, as
    the interglacial hit, raising sea levels and cutting everyone off from
    each other.
    The key here is Aquatic Ape.
    The most important piece, the one population that is common to all,
    that is everyone's ancestor is that Waterside population! They were
    the conduit. As they encountered these far flung groups, interbreeding, exchanging DNA (acquiring advantageous adaptations) they carried
    them along to the next group they encountered... and the next...
    We only say "Out of Africa" because of Toba. If Yellowstone explodes
    tomorrow it'll be like 50 Krakatoas exploding all at once. When Toba
    erupted it was roughly 2.8x Yellowstone! https://jtem.tumblr.com/image/176437002088
    (Not my image. I stole it off the web. But anyone can Google it if they
    doubt what I'm saying)
    THIS is why we say "Out of Africa."
    Toba exploded, Ground Zero was Sundaland -- the "Asia" in the "Out of
    Asia" scenario -- so the other major population center situated along
    the equator, away from where the worst of it was going to be, was
    Africa.
    There. That's the model. That's the fusion of Multiregionalism/Regional Continuity, Punctuated Equilibrium, Coastal Dispersal: Aquatic Ape. Lucy
    is an expected (predicted) biproduct of all that. She descended from a
    group that peeled off of an earlier group, pushed inland and adapted to
    a new environment... resulting in speciation.

    Our most-aquatic phase (H.erectus AFAWK) was probably early-Pleistocene.

    I think you're too conservative. I'd say that erectus is it. "She sang."

    AFAWK, erectus was our most-aquatic known ancestor:
    pachyosteosclerosis He>Hn>Hs (or, if you want, He>Hsn>Hss),
    but brain size Hn>Hs>He (evolution = slow?).

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  • From JTEM is so reasonable@21:1/5 to littor...@gmail.com on Wed Dec 14 17:38:04 2022
    littor...@gmail.com wrote:

    1) Lucy was at most only a partially "aquatic" ape, more precisely, it was an *aquarboreal* ape,
    something like (google) "wading gorilla". Ape ancestors have alway remained partly arboreal.

    "Vestigial."

    The problem here is that the observations support a number of conclusions.

    They are consistent with Lucy being descended from an aquatic population, and retaining unnecessary/vestigial traits.

    IMO, Miocene Hominoidea spread aquarboreally along Tethys Ocean (later Ind.Ocean + Med.Sea).

    Well they had to start somewhere, right? So that makes sense.

    2) The more aquatic (shallow-diving) Homo evolved later, late-Pliocene or possibly only early-Pleist.:
    Pleist.Homo reached overseas islands, evolved pachyosteosclerosis, large brains, platypelloidy etc.etc.

    Well Lucy is later.

    Yes, Gorilla & HP split c 8 Ma, when the northern part of the E.Afr.Rift formed (Red Sea),
    Lucy's ancestors followed the Rift ->Gorilla.
    Much later, Taung's ancestors followed the southern part of the E.Afr.Rift ->Pan.
    Gorilla & Pan evolved in //
    - late-Pliocene "gracile" afarensis//africanus,
    - early-Pleist. "robust" boisei//robustus,
    - today Afr.apes gorillas//chimps-bonobos.

    So our models fit. You're just getting into some firm details which I can't
    say are firm.

    I'd think Homo & Pan split when the Red Sea opened into the Ind.Ocean:

    Possible. If so, might've taken as much as a million years for Pan to evolve.

    -- Pan went right = E.Afr.coastal forests initially,
    -- Homo went left = S.Asian coastal forests initially ->shallow-diving early-Pleist.H.erectus Java etc.

    I'd like to explore this more. I do favor an origin of apes outside of Africa.

    *If* there ever was an "Out-of-Africa", it was late-Pleistocene, and perhaps not even all H.sapiens.

    I see it as < 80k years ago, after Toba.

    It's not that "Modern Man" evolved in Africa and then immediately decided
    that they hate the place, deciding to walk to Australia. Toba erupted with the energy of 250,000 nuclear warheads, or more, killing off most of Homo, and
    the populations best suited to survive were in Africa, and the one best suited to recover quickly & fill the vacuum was the sexually selected group...

    Which is one reason why I hate the words "Out of Africa." It doesn't tell us anything about human origins per se. I don't even think that the African
    group was that distinct. They were probably much better grouped with
    Eurasians, genetically, than populations elsewhere in Africa. The Bantu certainly were.




    -- --

    https://jtem.tumblr.com/post/703468006592905216

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  • From littoral.homo@gmail.com@21:1/5 to All on Thu Dec 15 02:40:24 2022
    Op donderdag 15 december 2022 om 02:38:05 UTC+1 schreef JTEM is so reasonable:
    littor...@gmail.com wrote:

    1) Lucy was at most only a partially "aquatic" ape, more precisely, it was an *aquarboreal* ape,
    something like (google) "wading gorilla". Ape ancestors have alway remained partly arboreal.

    "Lucy's skeleton, which is 40 % complete, was recovered in Ethiopia in what was an ancient lake near fossilized remains of crocodiles, turtle eggs and crab claws."
    https://www.cbc.ca/news/science/lucy-hominin-death-1.3739951

    "Vestigial."
    The problem here is that the observations support a number of conclusions. They are consistent with Lucy being descended from an aquatic population, and retaining unnecessary/vestigial traits.

    Lucy lived in swamp forests, : aquarboreal (aqua=water, arbor=tree):
    wading bipedally between the trees & climbing arms overhead,
    google "gorilla wading", e.g. in Ndoki.
    Lucy was no human ancestor, of course, but a Pliocene relative of Gorilla.

    IMO, Miocene Hominoidea spread aquarboreally along Tethys Ocean (later Ind.Ocean + Med.Sea).

    Well they had to start somewhere, right? So that makes sense.

    Yes, hylobatids live in SE.Asia, pongids live in SE.Asia, archaic Homo fossil lay on Java & even Flores...

    2) The more aquatic (shallow-diving) Homo evolved later, late-Pliocene or possibly only early-Pleist.:
    Pleist.Homo reached overseas islands, evolved pachyosteosclerosis, large brains, platypelloidy etc.etc.

    Well Lucy is later.

    No, no: Lucy c 3 Ma, H.erectus <2 Ma.

    Yes, Gorilla & HP split c 8 Ma, when the northern part of the E.Afr.Rift formed (Red Sea),
    Lucy's ancestors followed the Rift ->Gorilla.
    Much later, Taung's ancestors followed the southern part of the E.Afr.Rift ->Pan.
    Gorilla & Pan evolved in //
    - late-Pliocene "gracile" afarensis//africanus,
    - early-Pleist. "robust" boisei//robustus,
    - today Afr.apes gorillas//chimps-bonobos.

    So our models fit. You're just getting into some firm details which I can't say are firm.

    Pachyosteosclerosis (only Pleist.) is very "firm" (literally).

    I'd think Homo & Pan split when the Red Sea opened into the Ind.Ocean:

    Possible. If so, might've taken as much as a million years for Pan to evolve.

    -- Pan went right = E.Afr.coastal forests initially,
    -- Homo went left = S.Asian coastal forests initially ->shallow-diving early-Pleist.H.erectus Java etc.

    I'd like to explore this more. I do favor an origin of apes outside of Africa.

    Yes, IMO, c 30 Ma, the earliest Hominoidea reached the archipels then (full of coastal forests) between India & S.Eurasia.
    Late-Miocene hominids much later apparently lived along the Red Sea (we can call this Asia as well as Africa).

    *If* there ever was an "Out-of-Africa", it was late-Pleistocene, and perhaps not even all H.sapiens.

    I see it as < 80k years ago, after Toba.

    Yes, possible.

    It's not that "Modern Man" evolved in Africa and then immediately decided that they hate the place, deciding to walk to Australia. Toba erupted with the
    energy of 250,000 nuclear warheads, or more, killing off most of Homo, and the populations best suited to survive were in Africa, and the one best suited
    to recover quickly & fill the vacuum was the sexually selected group...
    Which is one reason why I hate the words "Out of Africa." It doesn't tell us anything about human origins per se. I don't even think that the African group was that distinct. They were probably much better grouped with Eurasians, genetically, than populations elsewhere in Africa. The Bantu certainly were.

    Yes, "OoA" is a very misleading slogan.

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  • From JTEM is so reasonable@21:1/5 to littor...@gmail.com on Thu Dec 15 13:50:26 2022
    littor...@gmail.com wrote:

    "Lucy's skeleton, which is 40 % complete, was recovered in Ethiopia in what was an
    ancient lake near fossilized remains of crocodiles, turtle eggs and crab claws."

    But other cites claims that she died from falling out of a tree! And regardless of manner of death, she wasn't lying there near any crocs, else she would have been a free meal. She was buried quickly. Or perhaps it was a low point in the water level, it was the fresh water equivalent to a "Tidal pool" left behind as the
    lake receded, perhaps anaerobic waters...

    I donno.

    "Vestigial."
    The problem here is that the observations support a number of conclusions. They are consistent with Lucy being descended from an aquatic population, and
    retaining unnecessary/vestigial traits.

    Lucy was no human ancestor, of course, but a Pliocene relative of Gorilla.

    I'm not at all concerned with whom Lucy's descendants were. I'm speaking of her ancestors. And her ancestors were aquatic. It wasn't a case of parallel evolution.
    Lucy is descended from a group that peeled off from the aquatic population, moved inland... possibly interbreeding with earlier groups too have done so, speeding
    it's evolution away from the Homo line.

    THAT is what I'm saying. I have no issue with Lucy giving rise to Pan or anything
    else.I'm focusing on who gave rise to her, and how.

    Well Lucy is later.

    No, no: Lucy c 3 Ma, H.erectus <2 Ma.

    Lucy came millions of years after the Miocene, which is where you want to put the split. So she's later than the split. She's Pliocene. Not Miocene.

    True, it could go the other way. She could be the product of a more primative population -- ardipithecus -- interbreeding with a more derived aquatic group, but it's six of one, half dozen of the other. You still need aquatic ape pushing
    inland leaving Lucy as a descendent.

    Again, I am focused on where Lucy came from, her ancestors, not what she
    led to, her descendants.






    -- --

    https://jtem.tumblr.com/post/703678017382285312

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  • From littoral.homo@gmail.com@21:1/5 to All on Fri Dec 16 02:58:28 2022
    Op donderdag 15 december 2022 om 22:50:27 UTC+1 schreef JTEM is so reasonable:

    "Lucy's skeleton, which is 40 % complete, was recovered in Ethiopia in what was an
    ancient lake near fossilized remains of crocodiles, turtle eggs and crab claws."

    But other cites claims that she died from falling out of a tree! And regardless
    of manner of death, she wasn't lying there near any crocs, else she would have
    been a free meal. She was buried quickly. Or perhaps it was a low point in the
    water level, it was the fresh water equivalent to a "Tidal pool" left behind as the
    lake receded, perhaps anaerobic waters... I donno.

    Yes. In any case, not in dry savanna.

    "Vestigial."
    The problem here is that the observations support a number of conclusions.
    They are consistent with Lucy being descended from an aquatic population, and
    retaining unnecessary/vestigial traits.

    Lucy was no human ancestor, of course, but a Pliocene relative of Gorilla.

    I'm not at all concerned with whom Lucy's descendants were. I'm speaking of her
    ancestors. And her ancestors were aquatic. It wasn't a case of parallel evolution.
    Lucy is descended from a group that peeled off from the aquatic population, moved inland... possibly interbreeding with earlier groups too have done so, speeding
    it's evolution away from the Homo line.
    THAT is what I'm saying. I have no issue with Lucy giving rise to Pan or anything
    else.I'm focusing on who gave rise to her, and how.

    Lucy's ancestors & other Mio-Pliocene hominoids & apiths were not "aquatic" sensu diving etc.
    AFAWK, they were bipeally wading + climbing vertically = aquarboreal:
    tail loss, centrally-placed = vertical spine, larger body, very broad body (sternum, thorax, pelvis), lateral arm + leg movements...
    That they sometimes might have dived is not impossible, but so far we have 0 indications.

    Our aquatic sensu frequently-diving phase came later: Pleistocene Homo: H.erectus: larger brain, pachy-osteo-sclerosis, island colonizations, external nose...

    Well Lucy is later.

    No, no: Lucy c 3 Ma, H.erectus <2 Ma.

    Lucy came millions of years after the Miocene, which is where you want to put the split. So she's later than the split. She's Pliocene. Not Miocene.
    True, it could go the other way. She could be the product of a more primative population -- ardipithecus -- interbreeding with a more derived aquatic group,
    but it's six of one, half dozen of the other. You still need aquatic ape pushing
    inland leaving Lucy as a descendent.
    Again, I am focused on where Lucy came from, her ancestors, not what she
    led to, her descendants.

    Again: Lucy & other apiths & Mio-Pliocene Hominoidea were aquarboreal
    = vertical waders-climbers in swamp forests.

    What probably happened (cf hylobatids & pongids & H.erectus in SE.Asia):
    Plate Tectonics (as described in my book p.299-300):

    1) 30 Ma India approached S.Asia = island archipel fm = coastal forests.
    The Catarrhini that reached these island forests became aquarboreal (aqua=water, arbor=tree).
    2) India underneath S.Asia split lesser (hylobatids) East & greater apes (pongids-hominids) West.
    Greater apes colonized Tethys(->Med.)Sea-coastal forests.
    3) The Mesopotamian Seaway closure 15 Ma split pongids-sivapiths East & hominids-dryopiths West.
    Pongids-sivapiths -> SE.Asia drove hylobatids higher into the trees.
    4) The E.Afr.Rift System after c 8 Ma split Red Sea hominids (Gorilla in EARS) from Homo-Pan.
    Gorilla-Praeanthropus e.g. late-Pliocene afarensis -> early-Pleistocene boisei etc.
    5) The Red Sea opening into the Gulf 5 Ma split Pan right (E.Afr.coastal forests) & Homo left (S.Asia).
    Pan-Australopithecus s.s. followed the southern EARS, e.g. africanus -> robustus (Pan // Gorilla).
    6) Homo (Ind.Ocean->Med.->Atl.coasts) became littoral: shallow-diving for shellfish etc.
    Initially seasonally inland along rivers. After 80 ka (enzymes MC->LC-PUFAs) independent from water.

    Simple, no? :-)

    1-5) aquarboreal Hominoidea
    6) littoral archaic Homo

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