The taxonomic attribution of African hominin postcrania from the Miocene through the Pleistocene:
associations and assumptions
Frederick Grine, Carrie S Mongle, John G Fleagle & Ashley S Hammond cs 2022 J.hum.Evol.173,103255 doi 10.1016/j.jhevol.2022.103255
Postcranial bones may provide valuable information about fossil taxa: locomotor habits, manipulative abilities, body sizes.
Distinctive postcranial featuresare sometimes noted in species diagnoses. Numerous isolated postcranial fossils have become accepted by many workers as belonging to a particular species,
but it is worthwhile revisiting the evidence for each attribution, before including them in comparative samples cf. descriptions of new fossils, functional analyses in relation to particular taxa, or in evolutionary contexts.
Some workers eschew the taxonomic attribution of postcrania as being less important (or interesting) than interpreting their functional morphology,
but it is impossible to consider the evolution of functional anatomy in a taxonomic & phylogenetic vacuum.
There are 21 widely recognized hominin taxa that have been described from sites in Africa dated from late-Miocene to mid-Pleistocene,
postcranial elements have been attributed to 17 of these.
The bones that have been thus assigned range from many parts of a skeleton, to isolated elements,
but the extent to which postcranial material can be reliably attributed to a specific taxon varies considerably, from site to site, and species to species,
it is often the subject of considerable debate.
Here, we review the postcranial remains attributed to African hominin taxa from late-Miocene to mid- & late-Pleistocene,
we place these assignations into categories of reliability.
The catalog of attribution presented here may serve as a guide for making taxonomic decisions in the future.
_______
Very beautiful & interesting paper, but what is "hominin"?
"hominin" = fossil relative of humans rather than of chimps, bonobos or gorillas?
The authors assume anthropocentrically (like most paleo-anthropologists) that australopiths ("hominins") are closer relatives of Homo than of Pan or Gorilla.
That is statistically & otherwise impossible, of course
(e.g. Pongo has several fossil relatives, but for some curious reason, it's believed that Pan nor Gorilla have +-no fossil relatives).
In fact, a'piths were no fossils halfway African apes & humans (as traditionally, but anthropocentrically believed),
but were halfway the hominid LCA (Gorilla-Homo-Pan) & extant African apes:
we showed that
-East-African a'piths were closer relatives of Gorilla than of Homo or Pan, -South-African a'piths were closer relatives of Pan than of Homo or Gorilla: -E & S.African a'piths evolved in parallel, schematically from late-Pliocene "gracile" to early-Pleistocene "robust":
-- Gorilla fossil subgenus Praeanthropus afarensis cs --> boisei cs //
-- Pan fossil subgenus Australopithecus africanus cs --> robustus cs,
e.g.
1994 Hum.Evol.9:121-139 "Australopithecines: ancestors of the African apes?"
1996 Hum.Evol.11:35-41 "Morphological distance between australopithecine, human and ape skulls".
The taxonomic attribution of African hominin postcrania from the Miocene through the Pleistocene:
associations and assumptions
Frederick Grine, Carrie S Mongle, John G Fleagle & Ashley S Hammond cs 2022 >J.hum.Evol.173,103255 doi 10.1016/j.jhevol.2022.103255
Postcranial bones may provide valuable information about fossil taxa: locomotor habits, manipulative abilities, body sizes.
Distinctive postcranial featuresare sometimes noted in species diagnoses. >Numerous isolated postcranial fossils have become accepted by many workers as belonging to a particular species,
but it is worthwhile revisiting the evidence for each attribution, before including them in comparative samples cf. descriptions of new fossils, functional analyses in relation to particular taxa, or in evolutionary contexts.
Some workers eschew the taxonomic attribution of postcrania as being less important (or interesting) than interpreting their functional morphology,
but it is impossible to consider the evolution of functional anatomy in a taxonomic & phylogenetic vacuum.
There are 21 widely recognized hominin taxa that have been described from sites in Africa dated from late-Miocene to mid-Pleistocene,
postcranial elements have been attributed to 17 of these.
The bones that have been thus assigned range from many parts of a skeleton, to isolated elements,
but the extent to which postcranial material can be reliably attributed to a specific taxon varies considerably, from site to site, and species to species,
it is often the subject of considerable debate.
Here, we review the postcranial remains attributed to African hominin taxa from late-Miocene to mid- & late-Pleistocene,
we place these assignations into categories of reliability.
The catalog of attribution presented here may serve as a guide for making taxonomic decisions in the future.
_______
Very beautiful & interesting paper, but what is "hominin"?
"hominin" = fossil relative of humans rather than of chimps, bonobos or gorillas?
The authors assume anthropocentrically (like most paleo-anthropologists) that australopiths ("hominins") are closer relatives of Homo than of Pan or Gorilla.
That is statistically & otherwise impossible, of course
(e.g. Pongo has several fossil relatives, but for some curious reason, it's believed that Pan nor Gorilla have +-no fossil relatives).
In fact, a'piths were no fossils halfway African apes & humans (as traditionally, but anthropocentrically believed),
but were halfway the hominid LCA (Gorilla-Homo-Pan) & extant African apes:
we showed that
-East-African a'piths were closer relatives of Gorilla than of Homo or Pan, >-South-African a'piths were closer relatives of Pan than of Homo or Gorilla: >-E & S.African a'piths evolved in parallel, schematically from late-Pliocene "gracile" to early-Pleistocene "robust":
-- Gorilla fossil subgenus Praeanthropus afarensis cs --> boisei cs //
-- Pan fossil subgenus Australopithecus africanus cs --> robustus cs,
e.g.
1994 Hum.Evol.9:121-139 "Australopithecines: ancestors of the African apes?"
1996 Hum.Evol.11:35-41 "Morphological distance between australopithecine, human and ape skulls".
_______The taxonomic attribution of African hominin postcrania from the Miocene through the Pleistocene:
associations and assumptions
Frederick Grine, Carrie S Mongle, John G Fleagle & Ashley S Hammond cs 2022 >J.hum.Evol.173,103255 doi 10.1016/j.jhevol.2022.103255
Postcranial bones may provide valuable information about fossil taxa: locomotor habits, manipulative abilities, body sizes.
Distinctive postcranial featuresare sometimes noted in species diagnoses. >Numerous isolated postcranial fossils have become accepted by many workers as belonging to a particular species,
but it is worthwhile revisiting the evidence for each attribution, before including them in comparative samples cf. descriptions of new fossils, functional analyses in relation to particular taxa, or in evolutionary contexts.
Some workers eschew the taxonomic attribution of postcrania as being less important (or interesting) than interpreting their functional morphology,
but it is impossible to consider the evolution of functional anatomy in a taxonomic & phylogenetic vacuum.
There are 21 widely recognized hominin taxa that have been described from sites in Africa dated from late-Miocene to mid-Pleistocene,
postcranial elements have been attributed to 17 of these.
The bones that have been thus assigned range from many parts of a skeleton, to isolated elements,
but the extent to which postcranial material can be reliably attributed to a specific taxon varies considerably, from site to site, and species to species,
it is often the subject of considerable debate.
Here, we review the postcranial remains attributed to African hominin taxa from late-Miocene to mid- & late-Pleistocene,
we place these assignations into categories of reliability.
The catalog of attribution presented here may serve as a guide for making taxonomic decisions in the future.
Very beautiful & interesting paper, but what is "hominin"?
"hominin" = fossil relative of humans rather than of chimps, bonobos or gorillas?
Thus, Hominini (hominins) is the maximum clade containing Homo sapiens Linnaeus 1758 but not Pan (originally Simia) troglodytes Blumenbach 1775.
Op zaterdag 12 november 2022 om 15:11:25 UTC+1 schreef Pandora:
...
Thus, Hominini (hominins) is the maximum clade containing Homo sapiens
Linnaeus 1758 but not Pan (originally Simia) troglodytes Blumenbach 1775.
Of course, thanks, my boy, that confirms that apiths are no Hominini.
Only incredible imbeciles believe apiths are closer relatives of Homo than of Pan or Gorilla.
Thus, Hominini (hominins) is the maximum clade containing Homo sapiens
Linnaeus 1758 but not Pan (originally Simia) troglodytes Blumenbach 1775.
Of course, thanks, my boy, that confirms that apiths are no Hominini.
Only incredible imbeciles believe apiths are closer relatives of Homo than of Pan or Gorilla.
Did you bother to follow any of the links he posted? Here's one https://sci-hub.se/10.1016/j.jhevol.2019.03.006
Expanded character sampling underscores
phylogenetic stability of Ardipithecus ramidus as a basal hominin
Phylogenetic relationships among hominins provide a necessary
framework for assessing their evolution. Reconstructing these
relationships hinges on the strength of the character data analyzed.
The phylogenetic position of Ardipithecus ramidus is critical to understanding early hominin evolution, and while many accept that
it is most likely the sister taxon to all later hominins, others have
argued that Ar.ramidus was ancestral to Pan. Although the study by
Strait & Grine (2004) suggested the former, available evidence
permitted only 26% of characters in their matrix to be assessed for Ar.ramidus. Fossils described subsequently by Suwa, White cs
(2009) have enabled the number of characters that can
be coded for this species to be expanded to 78 % of the matrix. Here,
we incorporate these new character data to evaluate their impact on
the phylogenetic relationships of Ar.ramidus. Moreover, we have
further revised the Strait & Grine (2004) matrix as necessitated by additions to the hypodigms of other fossil taxa. This updated matrix
was analyzed using both parsimony & Bayesian techniques in a
sequence of 4 iterative steps to independently evaluate the impact
of matrix & expanded character revisions on tree topology. Despite
the new data & matrix revisions, tree topology has remained
remarkably stable. The addition of new cranio-dental material has
served to markedly strengthen the support for the placement of
Ar.ramidus as being derived relative to Sahelanthropus, and as the
sister taxon of all later hominins. These findings support the
phylogenetic hypothesis originally proposed by White cs
(1994). This updated matrix provides a basis for the assessment of additional extinct spp.
littor...@gmail.com wrote:
Op zaterdag 12 november 2022 om 15:11:25 UTC+1 schreef Pandora:
...
Thus, Hominini (hominins) is the maximum clade containing Homo sapiensOf course, thanks, my boy, that confirms that apiths are no Hominini.
Linnaeus 1758 but not Pan (originally Simia) troglodytes Blumenbach 1775. >>
Only incredible imbeciles believe apiths are closer relatives of Homo than of Pan or Gorilla.
Did you bother to follow any of the links he posted? Here's one
https://sci-hub.se/10.1016/j.jhevol.2019.03.006
Expanded character sampling underscores
phylogenetic stability of Ardipithecus ramidus as
a basal hominin
That Verhaegen qualifies them as "incredible imbeciles" says a lot
about his own disrespectful, resentful, unqualified personality.
That Verhaegen qualifies them as "incredible imbeciles" says a lot
about his own disrespectful, resentful, unqualified personality.
Op zondag 13 november 2022 om 12:45:11 UTC+1 schreef Pandora:
...
That Verhaegen qualifies them as "incredible imbeciles" says a lot
about his own disrespectful, resentful, unqualified personality.
My little little boy, I do indeed qualify people who believe their Pleistocene ancestors ran after antelopes as incredible imbeciles.
Grow up!
Op zondag 13 november 2022 om 12:45:11 UTC+1 schreef Pandora:
That Verhaegen qualifies them as "incredible imbeciles" says a lot
about his own disrespectful, resentful, unqualified personality.
Haven't you even read what I wrote??
I do respect serious descriptions of PAs (but not the ridiculous atelope running religion, of course):
Quotations on gorilla-like features in large East African australopith crania: > Incisal dental microwear in A.afarensis is most similar to that observed in Gorilla. Ryan & Johanson 1989.suture and well onto an uninflated glabella; and extremely convex inferolateral margins of the orbits such as found in some gorillas. Walker cs 1986.
The composite skull reconstructed mostly from A.L.333 specimens looked very much like a small female gorilla. Johanson & Edey 1981:351.
Other primitive [or advanced gorilla-like? MV] features found in KNM-WT 17000, but not know or much discussed for A.afarensis, are: very small cranial capacity; low posterior profile of the calvaria; nasals extended far above the frontomaxillar
As for the maximum parietal breadth and the biauriculare in O.H.5 & KNM-ER 406 the robust australopithecines have values near the Gorilla mean: both the pongids & the robust australopithecines have highly pneumatized bases. Kennedy 1991.1981:74-75.
In O.H.5, the curious & characteristic features of the Paranthropus skull... parallel some of those of the gorilla. Robinson 1960.
The A.boisei lineage has been characterized by sexual dimorphism of the degree seen in modern Gorilla for the length of its known history. Leakey & Walker, 1988.
A.boisei teeth showed a relative absence of prism decussation; among extant hominoids, Gorilla enamel showed relatively little decussation .... Beynon & Wood, 1986 (cf. Beynon et al., 1991).
Quotations on chimp-like features in South African australopith crania:
Alan [Walker] has analysed a number of Au.robustus teeth and they fall into the fruit-eating category: their teeth patterns look like those of chimpanzees ... Then, when be looked at some H.erectus teeth, be found that the pattern changed. Leakey
The keystone nasal bone arrangement suggested as a derived diagnostic of Paranthropus [robustus] is found in an appreciable number of pongids, particularly clearly in some chimpanzees. Eckhardt 1987.chimpanzees, and (b) the curves for brain volume relative to body weight are essentially parallel in pongids & australopithecines, leading Hofman to conclude that as with pongids, the australopithecines probably differed only in size, not in design.
P.paniscus provides a suitable comparison for Australopithecus [Sts.5]; they are similar in body size, postcranial dimensions and ... even in cranial & facial features. Zihlman cs 1978.
A.africanus Sts.5, which... falls well within the range of Pan troglodytes, is markedly prognathous or hyperprognathous". Ferguson 1989.
In Taung, I see nothing in the orbits, nasal bones, and canine teeth definitely nearer to the human condition than the corresponding parts of the skull of a modern young chimpanzee. Woodward, 1925.
The Taung juvenile seems to resemble a young chimpanzee more closely than it resembles L338y-6, a juvenile A.boisei. Rak & Howell 1978.
In addition to similarities in facial remodeling it appears that Taung & Australopithecus in general, had maturation periods similar to those of the extant chimpanzee. Bromage 1985.
I estimate an adult capacity for Taung ranging from 404-420 cm2, with a mean of 412 cm2. Application of Passinghams curve for brain development in Pan is preferable to that for humans because (a) brain size of early hominids approximates that of
In Taung, pneumatization has also extended into the zygoma & hard palate. This is intriguing because an intrapalatal extension of the maxillary sinus has only been reported in chimpanzees & robust australopithecines among higher primates. Bromage &Dean 1985.
That the fossil ape Australopithecus [Taung] is distinguished from all living apes by the... unfused nasal bones as claimed by Dart (1940), cannot be maintained in view of the very considerable number of cases of separate nasal bones among orang-utans & chimpanzees of ages corresponding to that of Australopithecus. Schultz 1941.
Not my words, but those of repected PAs...
littor...@gmail.com wrote:
Op zaterdag 12 november 2022 om 15:11:25 UTC+1 schreef Pandora:
...
Thus, Hominini (hominins) is the maximum clade containing Homo sapiensOf course, thanks, my boy, that confirms that apiths are no Hominini.
Linnaeus 1758 but not Pan (originally Simia) troglodytes Blumenbach 1775. >>
Only incredible imbeciles believe apiths are closer relatives of Homo than of Pan or Gorilla.
Did you bother to follow any of the links he posted? Here's one
https://sci-hub.se/10.1016/j.jhevol.2019.03.006
Expanded character sampling underscores
phylogenetic stability of Ardipithecus ramidus as
a basal hominin
That Verhaegen qualifies them as "incredible imbeciles" says a lot
about his own disrespectful, resentful, unqualified personality.
Haven't you even read what I wrote??
I do respect serious descriptions of PAs (but not the ridiculous atelope running religion, of course):
Liar, you said "Only incredible imbeciles believe apiths are closer relatives of Homo than of Pan or Gorilla.", while all Mongle et al did
was test that hypothesis with much more and better data and methods
than you had at your disposal in 1994/96.
suture and well onto an uninflated glabella; and extremely convex inferolateral margins of the orbits such as found in some gorillas”. Walker cs 1986.Quotations on gorilla-like features in large East African australopith crania:
• “Incisal dental microwear in A.afarensis is most similar to that observed in Gorilla”. Ryan & Johanson 1989.
• The composite skull reconstructed mostly from A.L.333 specimens “looked very much like a small female gorilla”. Johanson & Edey 1981:351.
• “Other primitive [or advanced gorilla-like? MV] features found in KNM-WT 17000, but not know or much discussed for A.afarensis, are: very small cranial capacity; low posterior profile of the calvaria; nasals extended far above the frontomaxillar
Leakey 1981:74-75.• As for the maximum parietal breadth and the biauriculare in O.H.5 & KNM-ER 406 “the robust australopithecines have values near the Gorilla mean: both the pongids & the robust australopithecines have highly pneumatized bases”. Kennedy 1991.
• In O.H.5, “the curious & characteristic features of the Paranthropus skull... parallel some of those of the gorilla”. Robinson 1960.
• The A.boisei “lineage has been characterized by sexual dimorphism of the degree seen in modern Gorilla for the length of its known history”. Leakey & Walker, 1988.
• A.boisei teeth showed “a relative absence of prism decussation”; among extant hominoids, “Gorilla enamel showed relatively little decussation ...”. Beynon & Wood, 1986 (cf. Beynon et al., 1991).
Quotations on chimp-like features in South African australopith crania: >• “Alan [Walker] has analysed a number of Au.robustus teeth and they fall into the fruit-eating category: their teeth patterns look like those of chimpanzees ... Then, when be looked at some H.erectus teeth, be found that the pattern changed”.
that of chimpanzees, and (b) the curves for brain volume relative to body weight are essentially parallel in pongids & australopithecines, leading Hofman to conclude that ‘as with pongids, the australopithecines probably differed only in size, not in• “The ‘keystone’ nasal bone arrangement suggested as a derived diagnostic of Paranthropus [robustus] is found in an appreciable number of pongids, particularly clearly in some chimpanzees”. Eckhardt 1987.
• “P.paniscus provides a suitable comparison for Australopithecus [Sts.5]; they are similar in body size, postcranial dimensions and ... even in cranial & facial features”. Zihlman cs 1978.
• “A.africanus Sts.5, which... falls well within the range of Pan troglodytes, is markedly prognathous or hyperprognathous”". Ferguson 1989.
• In Taung, “I see nothing in the orbits, nasal bones, and canine teeth definitely nearer to the human condition than the corresponding parts of the skull of a modern young chimpanzee”. Woodward, 1925.
• “The Taung juvenile seems to resemble a young chimpanzee more closely than it resembles L338y-6”, a juvenile A.boisei. Rak & Howell 1978.
• “In addition to similarities in facial remodeling it appears that Taung & Australopithecus in general, had maturation periods similar to those of the extant chimpanzee”. Bromage 1985.
• “I estimate an adult capacity for Taung ranging from 404-420 cm2, with a mean of 412 cm2. Application of Passingham’s curve for brain development in Pan is preferable to that for humans because (a) brain size of early hominids approximates
Bromage & Dean 1985.• In Taung, “pneumatization has also extended into the zygoma & hard palate. This is intriguing because an intrapalatal extension of the maxillary sinus has only been reported in chimpanzees & robust australopithecines among higher primates”.
among orang-utans & chimpanzees of ages corresponding to that of Australopithecus”. Schultz 1941.• “That the fossil ape Australopithecus [Taung] ‘is distinguished from all living apes by the... unfused nasal bones…’ as claimed by Dart (1940), cannot be maintained in view of the very considerable number of cases of separate nasal bones
Not my words, but those of repected PAs...
Selective, out-of-context quote mining i
It's obvious that Verhaegen isn't well-versed in modern phylogenetic systematics. He's stuck in the previous century with his 1994/96
papers.
Some kudu runner:
It's obvious that Verhaegen isn't well-versed in modern phylogenetic systematics. He's stuck in the previous century with his 1994/96:-DDD
papers.
Your only *argument*!
Grow up, my little little boy.
No, no, my boy, e.g.
1) I've read +-all PA papers after that time, and all confirm my view:
- E.Afr.apiths were fossil relatives of Gorilla,
- S.Afr.apiths, of Pan (IOW, they were indeed more closely related to us than to Gorilla - got it??).
2) I've (co)written a few papers since 1996, incl.2 books:
1997 R Bender, --, N Oser Anthropol Anz 55:1-14 "Der Erwerb menschlicher Bipedie aus der Sicht der Aquatic Ape Theory"
1997 New Scientist 2091:53 "Sweaty humans"
1997 Hadewijch Antwerp 220pp "In den Beginne was het Water – Nieuwste Inzichten in de Evolutie van de Mens"
1998 MA Raath ... PV Tobias eds 1998 Dual Congress Univ Witwatersrand Jo'burg:128-9 "Australopithecine ancestors of African apes?"
1998 ibid.:131 "Human/ape brain differences and speech origins"
1998 --, P-F Puech ibid.:47 "Wetland apes: hominid palaeo-environment and diet"
1999 --, N McPhail, S Munro ESS Newsletter 50:4-12 "Bipedalism in chimpanzee and gorilla forebears"
1999 --, S Munro Water & Human Evolution Symposium Univ.Gent :11-23 "Australopiths wading? Homo diving?"
1999 --, S Munro Mother Tongue V:161-168 "Bipeds, tools and speech"
2000 --, P-F Puech Hum Evol 15:175-186 "Hominid lifestyle and diet reconsidered: paleo-environmental and comparative data"
2000 --, S Munro in J-L Dessalles, L Ghadakpour eds 2000 "The Evolution of Language" Ecole Nat Sup Télécommunications Paris:236-240 "The origins of phonetic abilities: a study of the comparative data with reference to the aquatic theory"
2002 --, S Munro Nutr Health 16:25-27 "The continental shelf hypothesis" 2002 --, P-F Puech, S Munro Trends Ecol Evol 17:212-7 (google: aquarboreal) "Aquarboreal ancestors?"
2004 --, S Munro Hum Evol 19:53-70 "Possible preadaptations to speech – a preliminary comparative approach"
2007 --, S Munro in SI Muñoz ed 2007 "Ecology Research Progress" Nova NY:1-4 "New directions in palaeoanthropology"
2007 --, S Munro, M Vaneechoutte, R Bender, N Oser ibid.:155-186 google econiche Homo "The original econiche of the genus Homo: open plain or waterside?"
2009 --, S Munro in NI Xirotiris cs eds 2009 "Fish and Seafood – Anthropological and Nutritional Perspectives" 28th ICAF Conference Kamilari:37-38 "Littoral diets in early hominoids and/or early Homo?"
2009 S Munro, -- ibid.:28-29 "Pachyosteosclerosis suggests archaic Homo exploited sessile littoral foods"
2010 New Scientist 2782:69 Lastword 16.10.10 "Oi, big nose!"
2011 --, S Munro HOMO – J compar hum Biol 62:237-247 "Pachyosteosclerosis suggests archaic Homo frequently collected sessile littoral foods"
2011 S Munro, -- in M Vaneechoutte cs eds 2011 ebook Bentham Sci Publ "Was Man More Aquatic in the Past?":82-105 "Pachyosteosclerosis in archaic Homo: heavy skulls for diving, heavy legs for wading?"
2011 --, S Munro, P-F Puech, M Vaneechoutte ibid.:67-81 "Early Hominoids: orthograde aquarboreals in flooded forests?"
2011 M Vaneechoutte, S Munro, -- ibid.:181-9 "Seafood, diving, song and speech"
2012 M Vaneechoutte, S Munro, -- HOMO J compar hum Biol 63:496-503 "Book review: Reply to John Langdon’s review of the eBook Was Man More Aquatic in the Past? Bentham Sci Publ"
2013 Hum Evol 28:237-266 "The aquatic ape evolves: common misconceptions and unproven assumptions about the so-called Aquatic Ape Hypothesis"
2016 E Schagatay cs (google: Schagatay Brenna reply) "A reply to Alice Roberts and Mark Maslin: Our ancestors may indeed have evolved at the shoreline – and here is why..."
2022 Eburon Acad Uitg Utrecht 325pp "De Evolutie van de Mens – Waarom wij rechtop lopen en kunnen spreken"
3) Recent insight, very interestingly IMO (preliminarily described in my recent book):
the connection between hominoid+hominid speciations & plate tectonics: -cercopith/hominoid split c 30 Ma: India approaching Eurasia: archipel fm: aquarborealism
-lesser/great ape split c 20 Ma: India fully under Eurasia -hominid-dryopith/pongid-sivapith split c 15 Ma: Mesopotamian Seaway closure -hominid s.l./s.s. split c 10 Ma: Red Sea colonisation
-HP/G split c 8 Ma: E.Afr.Rift fm: Gorilla-Praeanthropus -Homo/Pan-Australopith.s.s. split c 5 Ma: Red Sea opening into Ind.Ocean
Some kudu runner:
It's obvious that Verhaegen isn't well-versed in modern phylogenetic
systematics. He's stuck in the previous century with his 1994/96
papers.
:-DDD
Your only *argument*!
Grow up, my little little boy.
No, no, my boy, e.g.
1) I've read +-all PA papers after that time, and all confirm my view:
2) I've (co)written a few papers since 1996, incl.2 books:
2022 Eburon Acad Uitg Utrecht 325pp "De Evolutie van de Mens Waarom wij rechtop lopen en kunnen spreken"
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